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Genus Onthophagus

Insecta subclass Pterygota infraclass Neoptera superorder Holometabola order Coleoptera suborder Polyphaga infraorder Scarabeiformia superfamily Scarabaeoidea family Scarabaeidae subfamily Scarabaeinae tribe Onthophagini → genus Onthophagus Latreille 1802

urn:lsid:insecta.pro:taxonomy:830341

Onthophagus

Daughter taxa

Amphionthophagus Martin-Piera & Zunino, 1983 [subgenus]

Digitonthophagus Balthasar, 1959 [subgenus]

Onthophagus gazella

Furconthophagus Zunino, 1979 [subgenus]

Palaeonthophagus Zunino, 1979 [subgenus]

Onthophagus nuchicornis

Relictonthophagus Kabakov, 1979 [subgenus]

Trichonthophagus Zunino, 1979 [subgenus]

Onthophagus abacus Boucomont, 1921 [species]

Onthophagus abas Balthasar, 1946 [species]

Onthophagus abeillei D'orbigny, 1897 [species]

Onthophagus abreui Arrow, 1931 [species]

Onthophagus abruptus D'orbigny, 1913 [species]

Onthophagus absyrtus Balthasar, 1946 [species]

Onthophagus abyssinicus Gillet, 1925 [species]

Onthophagus academus Balthasar, 1946 [species]

Onthophagus accedens Lansberge, 1883 [species]

Onthophagus acerus Gillet, 1930 [species]

Onthophagus aciculatus Fahraeus, 1857 [species]

Onthophagus acrisius Balthasar, 1946 [species]

Onthophagus acuminatus Harold, 1880 [species]

Onthophagus acuticollis Gillet, 1927 [species]

O. a. sakishimanus

Onthophagus acuticornis Endrödi, 1973 [species]

Onthophagus acutus D'orbigny, 1908 [species]

Onthophagus adelaidae Hope, 1846 [species]

Onthophagus adelphus Gillet, 1930 [species]

Onthophagus admetus Balthasar, 1946 [species]

Onthophagus adornatus D'orbigny, 1904 [species]

Onthophagus adspersus D'orbigny, 1908 [species]

Onthophagus aegrotus Balthasar, 1967 [species]

Onthophagus aemulus Gillet, 1930 [species]

Onthophagus aeneoniger D'orbigny, 1913 [species]

Onthophagus aeneopiceus D'orbigny, 1902 [species]

Onthophagus aenescens Wiedemann, 1823 [species]

Onthophagus aequatus Péringuey, 1901 [species]

Onthophagus aequepubens D'orbigny, 1905 [species]

Onthophagus aerarius Reitter, 1892 [species]

Onthophagus aereidorsis D'orbigny, 1902 [species]

Onthophagus aeremicans D'orbigny, 1904 [species]

Onthophagus aereomaculatus Boucomont, 1914 [species]

Onthophagus aereopictus Boucomont, 1914 [species]

O. a. rubropictus

Onthophagus aerestriatus D'orbigny, 1913 [species]

Onthophagus aeruginosus Roth, 1851 [species]

Onthophagus aerumnosus Balthasar, 1963 [species]

Onthophagus aesopus Lansberge, 1882 [species]

Onthophagus aethiopicus D'orbigny, 1902 [species]

Onthophagus affinis Gillet, 1930 [species]

Onthophagus afghanus Petrovitz, 1961 [species]

Onthophagus africanus Lansberge, 1886 [species]

Onthophagus agaricophilus Arrow, 1931 [species]

Onthophagus agnus Gillet, 1925 [species]

Onthophagus ahenicollis D'orbigny, 1902 [species]

Onthophagus ahenomicans D'orbigny, 1902 [species]

Onthophagus akhaus Masumoto, Ochi & Hanboonsong, 2007 [species]

Onthophagus akinini Koenig, 1889 [species]

Onthophagus albarracinus Baraud, 1979 [species]

Onthophagus albicapillus D'Orbigny, 1902 [species]

Onthophagus albicomus D'orbigny, 1908 [species]

Onthophagus albicornis Palisot de Beauvois, 1805 [species]

Onthophagus albipennis Péringuey, 1908 [species]

Onthophagus albipodex D'orbigny, 1902 [species]

Onthophagus alcedo d'Orbigny 1913 [species]

Onthophagus alcyon Klug, 1855 [species]

Onthophagus alcyonides d'Orbigny 1913 [species]

Onthophagus aleppensis Redtenbacher, 1843 [species]

O. a. hyrcanicus

Onthophagus alexeevi Tarasov, Krikken & Huijbregts, 2010 [species]

Onthophagus alfuricus Huijbregts & Krikken, 2012 [species]

Onthophagus alienus Frey, 1958 [species]

Onthophagus allojavanus Huijbregts & Krikken, 2011 [species]

Onthophagus alluaudi D'orbigny, 1902 [species]

Onthophagus aloysiellus Zunino, 1977 [species]

Onthophagus alquirta Matthews, 1972 [species]

Onthophagus alternans Raffray, 1877 [species]

Onthophagus altidorsis D'orbigny, 1905 [species]

Onthophagus altilamina D'orbigny, 1905 [species]

Onthophagus altivagans Howden & Génier, 2004 [species]

Onthophagus amamiensis Nomura, 1965 [species]

Onthophagus ambang Krikken & Huijbregts, 2011 [species]

O. a. morowali

Onthophagus amblychromatus Marcus, 1917 [species]

Onthophagus amicus Gillet, 1925 [species]

Onthophagus amirus Kabakov, 1982 [species]

Onthophagus amoenus D'orbigny, 1908 [species]

Onthophagus amphicoma Boucomont, 1914 [species]

Onthophagus amphinasus Arrow, 1931 [species]

Onthophagus amphioxus Arrow, 1933 [species]

Onthophagus amplexus Sharp, 1875 [species]

Onthophagus amplipennis D'orbigny, 1905 [species]

Onthophagus anatolicus Petrovitz, 1962 [species]

Onthophagus anceyi Boucomont, 1921 [species]

Onthophagus anchommatus Lea, 1923 [species]

Onthophagus andalusicus Waltl, 1835 [species]

O. a. persianus

Onthophagus andersoni HOWDEN & GILL, 1987 [species]

Onthophagus andonarensis Boucomont, 1914 [species]

Onthophagus andrewdavisi Huijbregts & Krikken, 2009 [species]

Onthophagus andrewesi Arrow, 1931 [species]

Onthophagus androgynus D'orbigny, 1905 [species]

Onthophagus anewtoni Howden & Génier, 2004 [species]

Onthophagus angolensis Paulian, 1937 [species]

Onthophagus angorensis Petrovitz 1963 [species]

Onthophagus anguicorius Boucomont, 1914 [species]

Onthophagus angularis D'orbigny, 1908 [species]

Onthophagus anguliceps Boucomont, 1914 [species]

Onthophagus angulicornis D'orbigny, 1908 [species]

Onthophagus angustatus Boucomont, 1914 [species]

Onthophagus anisocerus Erichson, 1842 [species]

Onthophagus anitidus Ochi & Kon, 2005 [species]

Onthophagus annoyeri Moretto, 2010 [species]

Onthophagus anogeissii Cambefort, 1984 [species]

Onthophagus anomalicollis Frey, 1973 [species]

Onthophagus anomalipes D'orbigny, 1902 [species]

Onthophagus antennalis Frey, 1961 [species]

Onthophagus anthracinus Harold, 1873 [species]

Onthophagus antillarum Arrow, 1903 [species]

Onthophagus antilocapra Balthasar, 1937 [species]

Onthophagus antoinei Walter, 1990 [species]

Onthophagus aokii Nomura, 1976 [species]

Onthophagus aper Sharp, 1875 [species]

Onthophagus aphodioides Lansberge, 1883 [species]

Onthophagus apicetinctus D'orbigny, 1898 [species]

Onthophagus apiciosus D'orbigny, 1902 [species]

Onthophagus apilularius Masumoto, 1995 [species]

Onthophagus apterus Matthews, 1972 [species]

Onthophagus apunneea Masumoto, Ochi & Hanboonsong, 2007 [species]

Onthophagus arai Masumoto, 1989 [species]

Onthophagus arayai Ochi & Kon, 2007 [species]

Onthophagus arboreus Arrow, 1931 [species]

Onthophagus arcifer D'orbigny, 1904 [species]

Onthophagus ardoini Frey, 1971 [species]

Onthophagus areolatus D'orbigny, 1905 [species]

Onthophagus argyropygus Gillet, 1927 [species]

Onthophagus aries Kirby, 1825 [species]

Onthophagus arkoola STOREY & WEIR, 1990 [species]

Onthophagus armatus Blanchard, 1853 [species]

Onthophagus armicollis D'Orbigny, 1904 [species]

Onthophagus arnoldii Kabakov, 1982 [species]

Onthophagus arrilla Matthews, 1972 [species]

Onthophagus arunachalensis Biswas & Chatterjee, 1985 [species]

Onthophagus arunensis Scheuern, 1995 [species]

Onthophagus aschenborni Frey, 1975 [species]

Onthophagus ashanticola Frey, 1973 [species]

Onthophagus asiaticus Endrödi, 1973 [species]

Onthophagus asimilis Péringuey, 1901 [species]

Onthophagus asper Macleay, 1864 [species]

Onthophagus asperatus D'orbigny, 1905 [species]

Onthophagus aspericeps D'orbigny, 1908 [species]

Onthophagus aspericollis Lansberge, 1885 [species]

Onthophagus asperipennis D'orbigny, 1902 [species]

Onthophagus aspernatus D'orbigny, 1907 [species]

Onthophagus asperodorsatus HOWDEN & GILL, 1993 [species]

Onthophagus asperrimus D'orbigny, 1902 [species]

Onthophagus asperulus D'orbigny, 1905 [species]

Onthophagus astigma D'orbigny, 1904 [species]

Onthophagus ater Waterhouse, 1875 [species]

Onthophagus athiensis D'orbigny, 1913 [species]

Onthophagus atratus Kabakov, 1983 [species]

Onthophagus atricapillus D'Orbigny, 1908 [species]

Onthophagus atriclaviger d'Orbigny, 1905 [species]

Onthophagus atricolor D'orbigny, 1907 [species]

Onthophagus atridorsis D'orbigny, 1902 [species]

Onthophagus atriglabrus HOWDEN & GILL, 1987 [species]

Onthophagus atripennis Waterhouse, 1875 [species]

Onthophagus atroaereus D'orbigny, 1908 [species]

Onthophagus atrofasciatus D'orbigny, 1905 [species]

Onthophagus atronitidus D'orbigny, 1902 [species]

Onthophagus atropolitus D'Orbigny, 1902 [species]

Onthophagus atrosericeus Boucomont, 1932 [species]

Onthophagus atrosetosus D'Orbigny, 1902 [species]

Onthophagus atrostriatus D'orbigny, 1913 [species]

Onthophagus atrovirens D'orbigny, 1905 [species]

Onthophagus atrovittatus D'orbigny, 1908 [species]

Onthophagus atrox Harold, 1867 [species]

Onthophagus auratus Fabricius, 1801 [species]

Onthophagus aureiceps d'Orbigny, 1902 [species]

Onthophagus aureofuscus Bates, 1887 [species]

Onthophagus aureopilosus Boucomont, 1914 [species]

Onthophagus auriculatus Klug, 1855 [species]

Onthophagus aurifex Harold, 1877 [species]

Onthophagus auritus Erichson, 1842 [species]

Onthophagus australis Guérin-Méneville, 1830 [species]

Onthophagus avocetta Arrow, 1933 [species]

Onthophagus avocettoides Kabakov, 1983 [species]

Onthophagus axillaris Boheman, 1860 [species]

Onthophagus aztecus ZUNINO & HALFFTER, 1988 [species]

Onthophagus azusae Ochi & Kon, 2006 [species]

Onthophagus babaulti D'orbigny, 1915 [species]

Onthophagus babirussa Eschscholtz, 1822 [species]

Onthophagus badeni Harold, 1875 [species]

Onthophagus baenzigeri Masumoto, Ochi & Hanboonsong, 2007 [species]

Onthophagus baeri Boucomont, 1914 [species]

Onthophagus bakeri Boucomont, 1919 [species]

Onthophagus balawaicus Scheuern, 1995 [species]

Onthophagus baloghi Balthasar, 1967 [species]

Onthophagus balthasari Vsetecka, 1939 [species]

Onthophagus bambra Matthews, 1972 [species]

Onthophagus bandamai Cambefort, 1984 [species]

Onthophagus bangueyensis Boucomont, 1914 [species]

Onthophagus baoule Cambefort, 1984 [species]

Onthophagus baraudi Nicolas, 1964 [species]

Onthophagus barbieri Paulian, 1978 [species]

Onthophagus barretti Génier & Howden, 1999 [species]

Onthophagus barriorum Walter, 1990 [species]

Onthophagus bartosi Balthasar, 1966 [species]

Onthophagus basakata WALTER & CAMBEFORT, 1977 [species]

Onthophagus basicruentatus Goidanich, 1926 [species]

Onthophagus basilewskyi Frey, 1961 [species]

Onthophagus basipustulatus Heyden, 1889 [species]

Onthophagus bassariscus ZUNINO & HALFFTER, 1988 [species]

Onthophagus bateke WALTER & CAMBEFORT, 1977 [species]

Onthophagus batillifer Harold, 1875 [species]

Onthophagus batui Huijbregts & Krikken, 2009 [species]

Onthophagus bayeri Balthasar, 1942 [species]

Onthophagus baykanus Kabakov, 1994 [species]

Onthophagus bechynei Frey, 1953 [species]

Onthophagus beelarong STOREY & WEIR, 1990 [species]

Onthophagus beesoni Arrow, 1931 [species]

Onthophagus beiranus Péringuey, 1908 [species]

Onthophagus belinga Walter, 1989 [species]

Onthophagus bellus D'orbigny, 1905 [species]

Onthophagus belorhinus Bates, 1887 [species]

Onthophagus benedictorum WALTER & CAMBEFORT, 1977 [species]

Onthophagus bengalensis Harold, 1886 [species]

Onthophagus bengali Gordon & Oppenheimer, 1977 [species]

Onthophagus benguellianus Paulian, 1937 [species]

Onthophagus bennigseni Paulian, 1937 [species]

Onthophagus bequaerti D'orbigny, 1915 [species]

Onthophagus bergeri Frey, 1975 [species]

Onthophagus bernaudi Cambefort & nicolas, 1991 [species]

Onthophagus betschuanus Frey, 1975 [species]

Onthophagus biarmatus D'Orbigny, 1908 [species]

Onthophagus bicallifrons D'orbigny, 1902 [species]

Onthophagus bicallosus Klug, 1855 [species]

Onthophagus bicarinaticeps Lea, 1923 [species]

Onthophagus bicarinatus Gomes Alves, 1944 [species]

Onthophagus bicavicollis Lea, 1923 [species]

Onthophagus bicavifrons D'orbigny, 1902 [species]

Onthophagus bicolensis Ochi & Kon, 2006 [species]

O. b. asin

Onthophagus bicolor Raffray, 1877 [species]

Onthophagus biconifer D'orbigny, 1905 [species]

Onthophagus bicornis Macleay, 1888 [species]

Onthophagus bicristatus D'orbigny, 1905 [species]

Onthophagus bicristiger D'orbigny, 1913 [species]

Onthophagus bicuneus Kabakov, 2008 [species]

Onthophagus bidens Olivier, 1789 [species]

Onthophagus bidentatus Drapiez, 1819 [species]

Onthophagus bidentifrons D'orbigny, 1902 [species]

Onthophagus biexcavatus D'orbigny, 1898 [species]

Onthophagus bifasciatus Fabricius, 1781 [species]

Onthophagus bifidicornis D'orbigny, 1902 [species]

Onthophagus bifidus Reiche, 1849 [species]

Onthophagus bifrons D'orbigny, 1905 [species]

Onthophagus bilingula Balthasar, 1965 [species]

Onthophagus bimarginatus D'orbigny, 1902 [species]

Onthophagus bimetallicus D'orbigny, 1907 [species]

Onthophagus bindaree STOREY & WEIR, 1990 [species]

Onthophagus binodis Thunberg, 1818 [species]

Onthophagus binodosus D'orbigny, 1908 [species]

Onthophagus binodulus D'orbigny, 1913 [species]

Onthophagus binyana STOREY & WEIR, 1990 [species]

Onthophagus biplagiatus Thomson, 1858 [species]

Onthophagus birugatus D'orbigny, 1902 [species]

Onthophagus birugifer D'orbigny, 1908 [species]

Onthophagus bisbicornis D'orbigny, 1909 [species]

Onthophagus biscarinulatus Huijbregts & Krikken, 2012 [species]

Onthophagus bisectus Arrow, 1931 [species]

Onthophagus bisignatus D'orbigny, 1913 [species]

Onthophagus bison Boucomont, 1919 [species]

Onthophagus bistiniocelloides Krikken, 1986 [species]

Onthophagus bituber D'orbigny, 1904 [species]

Onthophagus bituberans D'orbigny, 1905 [species]

Onthophagus bituberculatus Olivier, 1789 [species]

Onthophagus bivertex Heyden, 1887 [species]

Onthophagus blackburni Shipp, 1895 [species]

Onthophagus blackwoodensis Blackburn, 1891 [species]

Onthophagus blanchardi Harold, 1869 [species]

Onthophagus blumei Lansberge, 1883 [species]

Onthophagus bocandei D'orbigny, 1904 [species]

Onthophagus bokiaunus Masumoto, 1995 [species]

Onthophagus bomberaianus Balthasar, 1969 [species]

Onthophagus bonengus Kabakov, 1994 [species]

Onthophagus bonorae Zunino, 1976 [species]

Onthophagus bonsae Zunino, 1976 [species]

Onthophagus boops D'orbigny, 1905 [species]

Onthophagus boranus Gestro, 1895 [species]

Onthophagus borassi Cambefort, 1984 [species]

Onthophagus borneensis Harold, 1877 [species]

Onthophagus bornemisszai Matthews, 1972 [species]

Onthophagus bornemisszanus Matthews, 1972 [species]

Onthophagus bottegi Gestro, 1895 [species]

Onthophagus boucomonti Paulian, 1931 [species]

Onthophagus bourgognei Paulian, 1945 [species]

Onthophagus bovinus Péringuey, 1892 [species]

Onthophagus brachypterus ZUNINO & HALFFTER, 1997 [species]

Onthophagus brazzavillianus Balthasar, 1974 [species]

Onthophagus brendelli Ochi, Kon & Barclay, 2008 [species]

Onthophagus breviceps D'orbigny, 1902 [species]

Onthophagus brevicollis Arrow, 1907 [species]

Onthophagus breviconus Génier & Howden, 1999 [species]

Onthophagus brevigena D'orbigny, 1902 [species]

Onthophagus brevipennis D'orbigny, 1902 [species]

Onthophagus brevisetis D'orbigny, 1915 [species]

Onthophagus brittoni Paulian, 1948 [species]

Onthophagus brivioi Frey, 1973 [species]

Onthophagus bronzeus Arrow, 1907 [species]

Onthophagus brooksi Matthews, 1972 [species]

Onthophagus brucei Reiche 1847 [species]

O. b. cyanostolus, O. b. njombeensis

Onthophagus brunellii Müller, 1942 [species]

Onthophagus brutus Arrow, 1931 [species]

Onthophagus bubalus Harold, 1867 [species]

Onthophagus buculus Mannerheim, 1829 [species]

Onthophagus bufulus Arrow, 1941 [species]

Onthophagus bunamin Matthews, 1972 [species]

Onthophagus bundara STOREY & WEIR, 1990 [species]

Onthophagus bundutuhanensis Ochi, Kon & Barclay, 2008 [species]

Onthophagus burchelli D'orbigny, 1908 [species]

Onthophagus busiris Balthasar, 1969 [species]

Onthophagus bytinskii Balthasar, 1960 [species]

Onthophagus caelator Balthasar, 1967 [species]

Onthophagus caesariatus Boucomont, 1921 [species]

Onthophagus caffrarius D'orbigny, 1902 [species]

Onthophagus calamophilus Krikken, 1977 [species]

Onthophagus calcaratus Boucomont, 1919 [species]

Onthophagus calchas Balthasar, 1967 [species]

Onthophagus calliger D'orbigny, 1913 [species]

Onthophagus callosipennis Boucomont, 1930 [species]

Onthophagus cambeforti Palestrini, 1992 [species]

Onthophagus cameloides D'Orbigny, 1898 [species]

Onthophagus camerunicus D'orbigny, 1905 [species]

Onthophagus cancer Lansberge, 1886 [species]

Onthophagus capella Kirby, 1818 [species]

Onthophagus capelliformis Gillet, 1925 [species]

Onthophagus capellinus Frey, 1963 [species]

Onthophagus capitatus Castelnau, 1840 [species]

Onthophagus capitosus Harold, 1867 [species]

Onthophagus caprai Frey, 1956 [species]

Onthophagus carayoni Cambefort, 1986 [species]

Onthophagus carcharias Harold, 1875 [species]

Onthophagus carinensis Boucomont, 1914 [species]

Onthophagus carinicollis Raffray, 1877 [species]

Onthophagus carinidorsis D'orbigny, 1908 [species]

Onthophagus carinifer D'orbigny, 1904 [species]

Onthophagus carinulatus Harold, 1877 [species]

Onthophagus carmodensis Blackburn, 1907 [species]

Onthophagus carpanetoi Pittino, 1982 [species]

Onthophagus carpophilus Pereira & Halffter, 1961 [species]

Onthophagus castetsi Lansberge, 1887 [species]

Onthophagus catenatus Lansberge, 1883 [species]

Onthophagus catharinensis Paulian, 1936 [species]

Onthophagus catta Fabricius, 1787 [species]

Onthophagus cavia Boucomont, 1914 [species]

Onthophagus caviceps Frey, 1962 [species]

Onthophagus cavidorsis d'Orbigny, 1902 [species]

Onthophagus cavifrons Harold, 1886 [species]

Onthophagus cavivertex D'orbigny, 1913 [species]

Onthophagus centricornis Fabricius, 1798 [species]

Onthophagus centurio Lansberge, 1885 [species]

Onthophagus cephalophi Cambefort, 1984 [species]

Onthophagus ceratotherium Janssens, 1953 [species]

Onthophagus cervenkai Kabakov, 2008 [species]

Onthophagus cervicapra Boucomont, 1914 [species]

Onthophagus cervus Fabricius, 1798 [species]

Onthophagus chaingraiensis Masumoto, Ochi & Hanboonsong, 2007 [species]

Onthophagus chaiyaphumensis Masumoto, Ochi & Hanboonsong, 2002 [species]

Onthophagus chalcostolus D'Orbigny, 1902 [species]

Onthophagus championi Bates, 1887 [species]

Onthophagus chandrai Ochi, 2007 [species]

Onthophagus changshouensis Zhang, 1997 [species]

Onthophagus chapaensis Kabakov, 1983 [species]

Onthophagus chebaicus Boucomont, 1919 [species]

Onthophagus cheesmanae Arrow, 1941 [species]

Onthophagus chepara Matthews, 1972 [species]

Onthophagus cheyi Ochi & Kon, 2006 [species]

Onthophagus chiapanecus ZUNINO & HALFFTER, 1988 [species]

Onthophagus chineicus Kabakov, 1998 [species]

Onthophagus chinensis Balthasar, 1952 [species]

Onthophagus chirindanus D'orbigny, 1913 [species]

Onthophagus chloroderus D'orbigny, 1905 [species]

Onthophagus chlorophanus D'orbigny, 1902 [species]

Onthophagus choanicus D'orbigny, 1904 [species]

Onthophagus chremes Balthasar, 1969 [species]

Onthophagus chryses Bates, 1887 [species]

Onthophagus chrysoderus D'orbigny, 1905 [species]

Onthophagus chrysopes Bates, 1888 [species]

Onthophagus chrysurus Arrow, 1931 [species]

Onthophagus chumphonensis Masumoto, Ochi & Hanboonsong, 2008 [species]

Onthophagus cicer Balthasar, 1964 [species]

Onthophagus cincticollis D'orbigny, 1902 [species]

Onthophagus cinctifrons D'orbigny, 1902 [species]

Onthophagus cinctipennis Quedenfeldt, 1884 [species]

Onthophagus cineraceus D'orbigny, 1902 [species]

Onthophagus circulator Reitter, 1891 [species]

Onthophagus circulifer Arrow, 1931 [species]

Onthophagus circumdatus D'orbigny, 1913 [species]

Onthophagus citreum Boucomont, 1919 [species]

Onthophagus civettae Cambefort, 1984 [species]

Onthophagus clavisetis D'orbigny, 1905 [species]

Onthophagus clermonti Paulian, 1931 [species]

Onthophagus clitellarius D'orbigny, 1908 [species]

Onthophagus clitellifer Reitter, 1894 [species]

Onthophagus clivimerus Huijbregts & Krikken, 2011 [species]

Onthophagus cludtsi Ochi, 2003 [species]

Onthophagus clusifrons D'orbigny, 1905 [species]

Onthophagus clypealis Lea, 1923 [species]

Onthophagus clypeatus Blanchard, 1846 [species]

Onthophagus coenobita Herbst, 1783 [species]

Onthophagus coeruleicollis Arrow, 1907 [species]

Onthophagus cognatus Boucomont, 1921 [species]

Onthophagus coiffaiti Frey, 1970 [species]

Onthophagus colasi Paulian, 1937 [species]

Onthophagus colffsi Lansberge, 1883 [species]

Onthophagus collaris Kabakov, 1983 [species]

Onthophagus collinsi Krikken & Huijbregts, 1987 [species]

Onthophagus colmanti D'Orbigny, 1908 [species]

Onthophagus columbianus Boucomont, 1932 [species]

Onthophagus comatulus D'orbigny, 1905 [species]

Onthophagus cometes Bates, 1888 [species]

Onthophagus commottoides Kabakov, 1998 [species]

Onthophagus comottoi Lansberge, 1885 [species]

Onthophagus compactus Arrow, 1933 [species]

Onthophagus comperei Blackburn, 1903 [species]

Onthophagus compositus Lea, 1923 [species]

Onthophagus compressus Guérin-Méneville, 1855 [species]

Onthophagus concavifrons D'orbigny, 1904 [species]

Onthophagus concolor Sharp, 1878 [species]

Onthophagus concursator Balthasar, 1960 [species]

Onthophagus confertus Péringuey, 1908 [species]

Onthophagus confluens D'orbigny, 1902 [species]

Onthophagus confossus D'Orbigny, 1915 [species]

Onthophagus conradsi Balthasar, 1937 [species]

Onthophagus consentaneus Harold, 1867 [species]

Onthophagus conspersus Reitter, 1893 [species]

Onthophagus conspicuus Macleay, 1864 [species]

Onthophagus contiguicornis D'orbigny, 1913 [species]

Onthophagus convexicollis Boheman, 1858 [species]

Onthophagus convexus D'orbigny, 1908 [species]

Onthophagus cooloola STOREY & WEIR, 1990 [species]

Onthophagus coomani Paulian, 1931 [species]

Onthophagus coorgensis Arrow, 1931 [species]

Onthophagus coprimorphoides Huijbregts & Krikken, 2012 [species]

Onthophagus coprimorphus Gillet, 1930 [species]

Onthophagus coptorhinodes Péringuey, 1901 [species]

Onthophagus coracinoides Kabakov, 1983 [species]

Onthophagus coracinus Boucomont, 1914 [species]

Onthophagus coriaceoumbrosus Kohlmann & Solis, 2001 [species]

Onthophagus corniculatus Reiche, 1847 [species]

Onthophagus corniculiger D'orbigny, 1913 [species]

Onthophagus cornifrons Thomson, 1858 [species]

Onthophagus cornutus Ferreira, 1971 [species]

Onthophagus coronatus D'orbigny, 1902 [species]

Onthophagus corrosus Bates, 1887 [species]

Onthophagus coscineus Bates, 1887 [species]

Onthophagus costatus D'orbigny, 1913 [species]

Onthophagus costifer Gillet, 1930 [species]

Onthophagus costiger D'orbigny, 1915 [species]

Onthophagus costilatus D'orbigny, 1908 [species]

Onthophagus costulicollis Frey, 1975 [species]

Onthophagus crantor Balthasar, 1967 [species]

Onthophagus crassicollis Boucomont, 1913 [species]

Onthophagus cratippus Balthasar, 1969 [species]

Onthophagus creber D'Orbigny, 1905 [species]

Onthophagus cribellum D'orbigny, 1902 [species]

Onthophagus cribratus Lansberge, 1883 [species]

Onthophagus cribripennis D'orbigny, 1902 [species]

Onthophagus criniger D'orbigny, 1904 [species]

Onthophagus crinitus Harold, 1869 [species]

O. c. panamensis

Onthophagus cristatus D'orbigny, 1905 [species]

Onthophagus croesulus Bates, 1888 [species]

Onthophagus crotchi Harold, 1871 [species]

Onthophagus crucenotatus D'orbigny, 1905 [species]

Onthophagus cruciatus Ménétriès, 1832 [species]

Onthophagus cruciger Macleay, 1888 [species]

Onthophagus cruentatus Klug, 1855 [species]

Onthophagus cryptodicranius Kohlmann & Solis, 2001 [species]

Onthophagus cryptogenus Boucomont, 1914 [species]

Onthophagus cuneus Boucomont, 1924 [species]

Onthophagus cuniculus Macleay, 1864 [species]

Onthophagus cupreiceps Arrow, 1907 [species]

Onthophagus cupreopastillatus Ochi & Kon, 2006 [species]

Onthophagus cupreovirens D'orbigny, 1904 [species]

Onthophagus cupreus Harold, 1880 [species]

Onthophagus cupricollis Péringuey, 1888 [species]

Onthophagus curtipilis D'orbigny, 1905 [species]

Onthophagus curtulus Frey, 1961 [species]

Onthophagus curvicarinatus Boucomont, 1914 [species]

Onthophagus curvicornis Latreille, 1812 [species]

Onthophagus curvifrons D'orbigny, 1906 [species]

Onthophagus curvilamina Müller, 1942 [species]

Onthophagus curvispina Reitter, 1893 [species]

Onthophagus cuspidatus d'Orbigny, 1913 [species]

Onthophagus cyaneiceps D'orbigny, 1913 [species]

Onthophagus cyaneoniger D'orbigny, 1902 [species]

Onthophagus cyaneus Kabakov, 2006 [species]

Onthophagus cyanochlorus D'orbigny, 1902 [species]

Onthophagus cyanopterus D'orbigny, 1905 [species]

Onthophagus cyclographus Bates, 1887 [species]

Onthophagus cyobioides Boucomont, 1921 [species]

Onthophagus dacatrai Pittino, 2004 [species]

Onthophagus dama Fabricius, 1798 [species]

Onthophagus damaki Masumoto, Hanboonsong & Ochi, 2002 [species]

Onthophagus dandalu Matthews, 1972 [species]

Onthophagus danumcupreus Krikken & Huijbregts, 2009 [species]

Onthophagus danumensis Ochi, Kon & Barclay, 2009 [species]

Onthophagus danumicus Huijbregts & Krikken, 2012 [species]

Onthophagus dapcauensis Boucomont, 1921 [species]

Onthophagus dapitanensis Boucomont, 1919 [species]

Onthophagus darlingtoni Matthews, 1972 [species]

Onthophagus davisi Frey, 1975 [species]

Onthophagus dayacus Boucomont, 1914 [species]

Onthophagus debilis D'orbigny, 1905 [species]

Onthophagus deccanensis Frey, 1962 [species]

Onthophagus decedens Péringuey, 1904 [species]

Onthophagus decens Balthasar, 1964 [species]

Onthophagus declivicollis D'orbigny, 1902 [species]

Onthophagus declivis Harold, 1869 [species]

Onthophagus decolor D'orbigny, 1902 [species]

Onthophagus decoratus D'orbigny, 1905 [species]

Onthophagus decorsei D'orbigny, 1908 [species]

Onthophagus dedecor Wallengren, 1881 [species]

Onthophagus deflexicollis Lansberge, 1883 [species]

Onthophagus deflexus D'orbigny, 1908 [species]

Onthophagus dejongi Krikken, 1986 [species]

Onthophagus delahayei Josso, 2011 [species]

Onthophagus delicatulus Raffray, 1877 [species]

Onthophagus delicatus D'orbigny, 1915 [species]

Onthophagus deliensis Lansberge, 1885 [species]

O. d. bawangicus

Onthophagus dellacasai Pittino & Mariani, 1981 [species]

Onthophagus delphinensis D'orbigny, 1913 [species]

Onthophagus demarzi Frey, 1959 [species]

Onthophagus demeyeri Moretto, 2009 [species]

Onthophagus densatus Frey, 1963 [species]

Onthophagus densepunctatus Frey, 1963 [species]

Onthophagus densipilis D'orbigny, 1902 [species]

Onthophagus denticollis Lansberge, 1883 [species]

Onthophagus denticornis Boucomont, 1914 [species]

Onthophagus denticulatus D'orbigny, 1902 [species]

Onthophagus denudatus D'orbigny, 1902 [species]

Onthophagus depilatus D'orbigny, 1913 [species]

Onthophagus depilis D'orbigny, 1902 [species]

Onthophagus deplanatus Lansberge, 1883 [species]

Onthophagus depressicollis Boucomont, 1914 [species]

Onthophagus depressifrons D'orbigny, 1915 [species]

Onthophagus depressipennis Cambefort, 1975 [species]

Onthophagus derasus D'orbigny, 1902 [species]

Onthophagus desaegeri Frey, 1961 [species]

Onthophagus desectus Macleay, 1871 [species]

Onthophagus deterrens Péringuey, 1901 [species]

Onthophagus devagiriensis Schoolmeesters & Thomas, 2006 [species]

Onthophagus devexicornis Gillet, 1930 [species]

Onthophagus devexus Macleay, 1888 [species]

Onthophagus dhanjuricus Frey, 1973 [species]

Onthophagus diabolicus Harold 1877 [species]

Onthophagus diadematus D'orbigny, 1902 [species]

Onthophagus dicax Balthasar, 1966 [species]

Onthophagus dicella Bates, 1888 [species]

Onthophagus dicranius Bates, 1887 [species]

Onthophagus dicranocerus Gillet, 1925 [species]

Onthophagus dicranoides Balthasar, 1939 [species]

Onthophagus difficilis Le Guillou, 1844 [species]

Onthophagus digitatus Arrow, 1931 [species]

Onthophagus dignus Gillet, 1930 [species]

Onthophagus dilutus D'orbigny, 1905 [species]

Onthophagus dinjeera Storey & Weir, 1990 [species]

Onthophagus dinoderus D'orbigny, 1913 [species]

Onthophagus discedens Sharp, 1875 [species]

Onthophagus discolor Hope, 1841 [species]

Onthophagus discovirens D'orbigny, 1902 [species]

Onthophagus discretus Péringuey, 1901 [species]

Onthophagus dispar Péringuey, 1901 [species]

Onthophagus dissentaneus Balthasar, 1964 [species]

Onthophagus distichus Roth, 1851 [species]

Onthophagus ditus Péringuey, 1901 [species]

Onthophagus diversiformis Boucomont, 1914 [species]

Onthophagus diversus Reiche, 1847 [species]

Onthophagus dives Harold, 1877 [species]

Onthophagus dlabolai Balthasar, 1964 [species]

Onthophagus dohertyi D'orbigny, 1902 [species]

Onthophagus doiinthanonensis Masumoto, 1989 [species]

Onthophagus doipuiensis Masumoto, 1989 [species]

Onthophagus doisuthepensis Masumoto, 1989 [species]

Onthophagus doitungensis Masumoto & Ochi & Hanboonsong, 2002 [species]

Onthophagus doriae Harold, 1877 [species]

Onthophagus dorsipilulus HOWDEN & GILL, 1987 [species]

Onthophagus dorsofasciatus Fairmaire, 1893 [species]

Onthophagus dorsosignatus D'Orbigny, 1898 [species]

Onthophagus dorsuosus D'orbigny, 1902 [species]

Onthophagus drescheri Paulian, 1939 [species]

Onthophagus dubernardi Boucomont, 1914 [species]

Onthophagus dubitabilis Howden & Génier, 2004 [species]

Onthophagus duboulayi Waterhouse, 1894 [species]

Onthophagus ducorpsi D'orbigny, 1913 [species]

Onthophagus dudleyi Cambefort, 1980 [species]

Onthophagus dummal Matthews, 1972 [species]

Onthophagus dunningi Harold, 1869 [species]

Onthophagus duporti Boucomont, 1914 [species]

Onthophagus duvivieri D'orbigny, 1904 [species]

Onthophagus dux Sharp, 1875 [species]

Onthophagus dynastoides Arrow, 1931 [species]

Onthophagus ebenicolor D'orbigny, 1902 [species]

Onthophagus ebenus Péringuey, 1888 [species]

Onthophagus eburneus D'orbigny, 1913 [species]

Onthophagus echinus Boucomont, 1914 [species]

Onthophagus ecopas Josso & Prévost, 2006 [species]

Onthophagus egenus Harold, 1877 [species]

Onthophagus egregius Arrow, 1907 [species]

Onthophagus egurianus Matsumura, 1938 [species]

Onthophagus elegans Klug, 1832 [species]

Onthophagus elgoni D'Orbigny, 1915 [species]

Onthophagus eliptaminus Balthasar, 1969 [species]

Onthophagus elongatus Frey, 1954 [species]

Onthophagus emarginatus Mulsant 1842 [species]

Onthophagus embersoni Masumoto, Hanboonsong & Ochi, 2002 [species]

Onthophagus embrikianus Paulian, 1936 [species]

Onthophagus emeritus Péringuey, 1901 [species]

Onthophagus endota Matthews, 1972 [species]

Onthophagus endroedianus WALTER & CAMBEFORT, 1977 [species]

Onthophagus endroedii Frey, 1973 [species]

Onthophagus ensifer Boucomont, 1914 [species]

Onthophagus enuguensis Frey, 1975 [species]

Onthophagus ephippioderus Arrow 1907 [species]

Onthophagus epilamprus Bates, 1888 [species]

Onthophagus erectinasus D'orbigny, 1902 [species]

Onthophagus erichsoni Hope, 1841 [species]

Onthophagus escalerai D'orbigny, 1902 [species]

Onthophagus eschscholtzi Boucomont, 1924 [species]

Onthophagus euchlorus D'Orbigny, 1902 [species]

Onthophagus eulaminicornis Pittino, 2006 [species]

Onthophagus eulophus Bates, 1887 [species]

Onthophagus euryceros Kabakov, 1998 [species]

Onthophagus euzeti WALTER & CAMBEFORT, 1977 [species]

Onthophagus evae Balthasar, 1944 [species]

Onthophagus evanidus Harold, 1869 [species]

Onthophagus exasperatus Gerstaecker, 1871 [species]

Onthophagus excisiceps D'orbigny, 1902 [species]

Onthophagus excisus Reiche & Saulcy, 1856 [species]

Onthophagus excubitor Ziani & Gudenzi, 2006 [species]

Onthophagus exiguus Raffray, 1877 [species]

Onthophagus exilis D'orbigny, 1913 [species]

Onthophagus expansicornis Bates, 1891 [species]

Onthophagus exquisitus Arrow, 1931 [species]

Onthophagus extensicollis D'orbigny, 1907 [species]

Onthophagus extensus Harold, 1878 [species]

Onthophagus fabricii Waterhouse, 1894 [species]

Onthophagus falcarius Balthasar, 1966 [species]

Onthophagus falcifer Harold, 1880 [species]

Onthophagus falculatus Boucomont, 1914 [species]

Onthophagus fallaciosus Raffray, 1877 [species]

Onthophagus fallax D'orbigny, 1913 [species]

Onthophagus falsivigilans Masumoto, 1995 [species]

Onthophagus falsus Gillet, 1925 [species]

Onthophagus falzonii Goidanich, 1926 [species]

Onthophagus fasciatus Boucomont, 1914 [species]

Onthophagus fasciculiger D'orbigny, 1902 [species]

Onthophagus fasciolatus Boucomont, 1924 [species]

Onthophagus fastosus D'Orbigny, 1913 [species]

Onthophagus favrei Boucomont, 1914 [species]

Onthophagus feai D'orbigny, 1905 [species]

Onthophagus felix Arrow, 1931 [species]

Onthophagus ferox Harold, 1867 [species]

Onthophagus ferrari Matthews, 1972 [species]

Onthophagus filicornis Harold, 1873 [species]

Onthophagus fimetarius Roth, 1851 [species]

Onthophagus finschi Harold, 1877 [species]

Onthophagus fissiceps Macleay, 1888 [species]

Onthophagus fissicornis Steven, 1809 [species]

Onthophagus fissinasus Fairmaire, 1895 [species]

Onthophagus fitiniensis Cambefort, 1984 [species]

Onthophagus flagrans Reitter, 1893 [species]

Onthophagus flavibasis D'orbigny, 1904 [species]

Onthophagus flaviclava D'orbigny, 1902 [species]

Onthophagus flavicornis Germar, 1824 [species]

Onthophagus flavimargo D'orbigny, 1902 [species]

Onthophagus flavipennis D'orbigny, 1905 [species]

Onthophagus flavoapicalis Lea, 1923 [species]

Onthophagus flavocinctus Frey, 1975 [species]

Onthophagus flavolimbatus Klug, 1855 [species]

Onthophagus flavomaculatus Gillet, 1930 [species]

Onthophagus flavorufus D'orbigny, 1904 [species]

Onthophagus fletcheri Blackburn, 1903 [species]

Onthophagus flexicollis D'Orbigny, 1904 [species]

Onthophagus flexicornis D'orbigny, 1902 [species]

Onthophagus flexifrons D'orbigny, 1908 [species]

Onthophagus fodiens Waterhouse, 1875 [species]

Onthophagus foedus Boucomont, 1914 [species]

Onthophagus foliaceus Lansberge, 1886 [species]

Onthophagus foliiceps Quedenfeldt, 1884 [species]

Onthophagus foraminosus D'orbigny, 1902 [species]

Onthophagus formaneki Reitter, 1897 [species]

Onthophagus formosanus Gillet, 1924 [species]

Onthophagus fortigibber Reitter, 1909 [species]

Onthophagus fossibasis D'orbigny, 1908 [species]

Onthophagus fossidorsis D'Orbigny, 1905 [species]

Onthophagus fossifrons D'orbigny, 1902 [species]

Onthophagus fossor Arrow, 1931 [species]

Onthophagus fossulatus D'orbigny, 1908 [species]

Onthophagus foulliouxi Cambefort, 1971 [species]

Onthophagus foveatus Frey, 1957 [species]

Onthophagus foveicollis Boucomont, 1930 [species]

Onthophagus fracticornis Preyssler 1790 [species]

Onthophagus fradei Gomes Alves, 1956 [species]

Onthophagus frankenbergeri Balthasar, 1933 [species]

Onthophagus frankrozendaali Huijbregts & Krikken, 2012 [species]

Onthophagus frenchi Blackburn, 1903 [species]

Onthophagus freyi Moretto, 2004 [species]

Onthophagus fritschi D'orbigny, 1902 [species]

Onthophagus frontalis Raffray, 1877 [species]

Onthophagus frugivorus Arrow 1931 [species]

Onthophagus fugitivus Péringuey, 1901 [species]

Onthophagus fujiii Ochi & Kon, 1995 [species]

Onthophagus fujiokai Ochi & Araya, 1996 [species]

Onthophagus fuliginosus Erichson, 1842 [species]

Onthophagus fulvocinctus D'orbigny, 1902 [species]

Onthophagus fulvus Sharp, 1875 [species]

Onthophagus fumatus D'Orbigny, 1913 [species]

Onthophagus funebris Boucomont, 1919 [species]

Onthophagus funestus Moretto & Génier, 2010 [species]

Onthophagus fungicola D'orbigny, 1908 [species]

Onthophagus furcaticeps Masters, 1886 [species]

Onthophagus furcatoides Lansberge, 1886 [species]

Onthophagus furcatus Fabricius, 1781 [species]

Onthophagus furciceps Marseul, 1869 [species]

Onthophagus furcicollis Arrow, 1931 [species]

Onthophagus furcifer Boheman, 1860 [species]

Onthophagus furcillifer Bates, 1891 [species]

Onthophagus furciramus D'Orbigny, 1902 [species]

Onthophagus furculifer D'orbigny, 1905 [species]

Onthophagus furculus Fabricius, 1798 [species]

Onthophagus fuscatus D'orbigny, 1908 [species]

Onthophagus fuscidorsis D ORBIGNY, 1902 [species]

Onthophagus fuscivestis D'orbigny, 1902 [species]

Onthophagus fuscopunctatus Fabricius, 1798 [species]

Onthophagus fuscostriatus Boucomont, 1914 [species]

Onthophagus gabonensis Walter, 1989 [species]

Onthophagus gaesatus Boucomont, 1925 [species]

Onthophagus gagates Hope, 1831 [species]

Onthophagus gagatinus Gillet, 1930 [species]

Onthophagus gagatoides Kabakov, 2006 [species]

Onthophagus gaillardi D'orbigny, 1911 [species]

Onthophagus gajo Krikken, 1977 [species]

Onthophagus galeatus Boucomont, 1919 [species]

Onthophagus ganalensis Gestro, 1895 [species]

Onthophagus gandju Matthews, 1972 [species]

Onthophagus gangeticus Gillet, 1925 [species]

Onthophagus gangloffi Josso & Prévost, 2001 [species]

Onthophagus gangulu Matthews, 1972 [species]

Onthophagus garambae Frey, 1961 [species]

Onthophagus gazellinus Bates, 1887 [species]

Onthophagus geelongensis Blackburn, 1891 [species]

Onthophagus gemellatus Raffray, 1877 [species]

Onthophagus geminatus D'orbigny, 1907 [species]

Onthophagus geminifrons D'orbigny, 1905 [species]

Onthophagus gemma Sharp, 1875 [species]

Onthophagus gemmatus Péringuey, 1901 [species]

Onthophagus genuinus Kohlmann & Solis, 2001 [species]

Onthophagus germanus Gillet, 1927 [species]

Onthophagus gerstaeckeri Harold, 1867 [species]

Onthophagus geryon Balthasar, 1969 [species]

Onthophagus gestroi Harold, 1877 [species]

Onthophagus ghanensis Balthasar, 1966 [species]

Onthophagus gibber D'orbigny, 1905 [species]

Onthophagus gibberosus D'orbigny, 1905 [species]

Onthophagus gibbicollis Lansberge, 1885 [species]

Onthophagus gibbidorsis D'orbigny, 1902 [species]

Onthophagus gibbifrons D'orbigny, 1902 [species]

Onthophagus gibbiramus D'Orbigny, 1902 [species]

Onthophagus gibbulus Pallas, 1781 [species]

Onthophagus gibbus D'orbigny, 1913 [species]

Onthophagus gibsoni Howden & Génier, 2004 [species]

Onthophagus gidju Matthews, 1972 [species]

Onthophagus gigantivigilans Masumoto, Hanboonsong & Ochi, 2002 [species]

Onthophagus gilli Delgado & Howden, 2000 [species]

Onthophagus ginyunensis Vsetecka, 1942 [species]

Onthophagus giraffa Hausmann, 1807 [species]

Onthophagus girardinae Walter, 1989 [species]

Onthophagus giuseppecarpanetoi Tagliaferri & Moretto, 2012 [species]

Onthophagus glabratus Hope, 1842 [species]

Onthophagus gladiator Arrow 1907 [species]

Onthophagus glasunovi Koshantschikov, 1894 [species]

Onthophagus glaucinus Gillet, 1930 [species]

Onthophagus gnu Frey, 1955 [species]

Onthophagus godarra STOREY & WEIR, 1990 [species]

Onthophagus gonipa Krikken & Huijbregts, 2011 [species]

Onthophagus gonopygus D'orbigny, 1902 [species]

Onthophagus gorochovi Kabakov, 1998 [species]

Onthophagus gorodinskii Kabakov, 2008 [species]

Onthophagus gorokae Paulian, 1972 [species]

Onthophagus gosoli Masumoto, 1989 [species]

Onthophagus gothicus Gillet, 1930 [species]

Onthophagus gracilipes Boucomont, 1914 [species]

Onthophagus gradivus Balthasar, 1966 [species]

Onthophagus grandidorsis D'orbigny, 1913 [species]

Onthophagus grandifrons D'orbigny, 1907 [species]

Onthophagus grandivigilans Masumoto, 1995 [species]

Onthophagus graniceps D'orbigny, 1908 [species]

Onthophagus granosus D'orbigny, 1913 [species]

Onthophagus granulatus Boheman, 1858 [species]

Onthophagus granulifer Harold, 1886 [species]

Onthophagus granulifrons Frey, 1961 [species]

Onthophagus granulipennis Lansberge, 1886 [species]

Onthophagus granulum D'orbigny, 1904 [species]

Onthophagus granum Lansberge, 1885 [species]

Onthophagus graphicus Wallengren, 1881 [species]

Onthophagus grassei Walter, 1989 [species]

Onthophagus grataehelenae Kohlmann & Solis, 2001 [species]

Onthophagus gratus Arrow, 1931 [species]

Onthophagus gravearmatus Balthasar, 1944 [species]

Onthophagus gravis Walker, 1858 [species]

Onthophagus gravoti D'orbigny, 1908 [species]

Onthophagus griseoaeneus Lansberge, 1885 [species]

Onthophagus griseosetosus Arrow, 1931 [species]

Onthophagus grossepunctatus Reitter 1905 [species]

Onthophagus guatemalensis Bates, 1887 [species]

Onthophagus gulmarri Matthews, 1972 [species]

Onthophagus gulo Arrow 1931 [species]

Onthophagus gunsalami Ochi & Kon, 2005 [species]

Onthophagus gurburra Storey & Weir, 1990 [species]

Onthophagus guttatus Boheman, 1860 [species]

Onthophagus guttiger D'orbigny, 1905 [species]

Onthophagus haafi Frey, 1963 [species]

Onthophagus haagi Harold, 1867 [species]

Onthophagus haematopus Harold, 1875 [species]

Onthophagus hageni Lansberge, 1883 [species]

Onthophagus hajimei Masumoto, 1984 [species]

Onthophagus halffteri Zunino, 1981 [species]

Onthophagus hamaticeps Arrow, 1931 [species]

Onthophagus hamaticornis Gillet, 1930 [species]

Onthophagus haroldi Ballion, 1870 [species]

Onthophagus harpax Fabricius, 1801 [species]

Onthophagus hartiniae Ochi & Kon, 2009 [species]

Onthophagus hastifer Lansberge, 1885 [species]

Onthophagus hayashii Masumoto, 1991 [species]

Onthophagus hayashimasaoi Ochi & Kon, 1999 [species]

Onthophagus helciatus Harold, 1871 [species]

Onthophagus helmsi Blackburn, 1903 [species]

Onthophagus hemichalceus D'orbigny, 1915 [species]

Onthophagus hemichlorus D'orbigny, 1915 [species]

Onthophagus hemicyanus D'Orbigny, 1902 [species]

Onthophagus hemipygus Frey, 1964 [species]

Onthophagus herciniformis Moretto, 2010 [species]

Onthophagus hericius D'orbigny, 1902 [species]

Onthophagus hermonensis Baraud 1982 [species]

Onthophagus herus Péringuey, 1901 [species]

Onthophagus heteroclitus D'orbigny, 1911 [species]

Onthophagus heterorrhinus Lansberge, 1885 [species]

Onthophagus heurni Gillet, 1930 [species]

Onthophagus heydeni Harold, 1875 [species]

Onthophagus heyrovskyi Vsetecka, 1943 [species]

Onthophagus hiabunicus Kabakov, 1998 [species]

Onthophagus hidakai Ochi & Kon, 1995 [species]

Onthophagus hielkemai Krikken & Huijbregts, 2011 [species]

Onthophagus hikidai Ochi & Kon, 2006 [species]

Onthophagus hilaridis Cambefort, 1984 [species]

Onthophagus hilaris D'orbigny, 1905 [species]

Onthophagus hildebrandti Harold, 1878 [species]

Onthophagus hindu Arrow 1931 [species]

Onthophagus hingstoni Arrow 1931 [species]

Onthophagus hippopotamus Harold, 1869 [species]

Onthophagus hirculus Mannerheim, 1829 [species]

Onthophagus hircus Billberg, 1815 [species]

Onthophagus hiroyukii Ochi, 2007 [species]

Onthophagus hirsutulus Lansberge, 1883 [species]

Onthophagus hirsutus D'orbigny, 1902 [species]

Onthophagus hirtellus Frey, 1957 [species]

Onthophagus hirticulus d'Orbigny 1915 [species]

Onthophagus hirtipodex D'orbigny, 1904 [species]

Onthophagus hirtuosus Gillet, 1930 [species]

Onthophagus hirtus Illiger, 1803 [species]

Onthophagus hispanicus Baraud, 1963 [species]

Onthophagus hissariensis Kabakov, 2006 [species]

Onthophagus histeriformis Boucomont, 1914 [species]

Onthophagus histrio D'orbigny, 1902 [species]

Onthophagus hoberlandti Balthasar, 1966 [species]

Onthophagus holosericeus Harold, 1877 [species]

Onthophagus holzi Boucomont, 1914 [species]

Onthophagus hoogstraali Saylor, 1943 [species]

Onthophagus hoplocerus Lea, 1923 [species]

Onthophagus hoplothorax Gillet, 1930 [species]

Onthophagus horii Ochi, Kon & Tsubaki, 2009 [species]

Onthophagus horni Balthasar, 1935 [species]

Onthophagus horrens D'orbigny, 1908 [species]

Onthophagus horribilis Balthasar, 1969 [species]

Onthophagus horridus D'orbigny, 1908 [species]

Onthophagus howdenorum ZUNINO & HALFFTER, 1988 [species]

Onthophagus hsui Masumoto, Chen & Ochi, 2004 [species]

Onthophagus hulstaerti Boucomont, 1932 [species]

Onthophagus humpatensis Lansberge, 1886 [species]

Onthophagus hyaena Fabricius, 1801 [species]

Onthophagus hyalcyon Palestrini & Giacone, 1989 [species]

Onthophagus hystrix Boucomont 1914 [species]

Onthophagus ibex Fabricius, 1792 [species]

Onthophagus ieti Cambefort, 1984 [species]

Onthophagus ifugaoensis Ochi & Kon, 2006 [species]

Onthophagus igneus Vigors, 1825 [species]

Onthophagus ignitus D'Orbigny, 1905 [species]

Onthophagus igualensis Bates, 1887 [species]

Onthophagus illotus Péringuey, 1901 [species]

Onthophagus illyricus Scopoli, 1763 [species]

Onthophagus imbellis D'orbigny, 1905 [species]

Onthophagus imberbis D'orbigny, 1902 [species]

Onthophagus imbutus Sharp, 1875 [species]

Onthophagus immundus Boheman, 1858 [species]

Onthophagus impar Kabakov, 2006 [species]

Onthophagus imperator Castelnau, 1840 [species]

Onthophagus importunus Péringuey, 1901 [species]

Onthophagus impressicollis Boheman, 1860 [species]

Onthophagus impunctatus D'orbigny, 1904 [species]

Onthophagus impurus Harold, 1868 [species]

Onthophagus inaequalis D'orbigny, 1902 [species]

Onthophagus incantatus Balthasar, 1967 [species]

Onthophagus incanus Macleay, 1888 [species]

Onthophagus incertus D'Orbigny, 1897 [species]

Onthophagus incisus Harold, 1877 [species]

Onthophagus includens D'orbigny, 1905 [species]

Onthophagus inclusus D'orbigny, 1905 [species]

Onthophagus incollaris Kabakov, 2008 [species]

Onthophagus incornutus Macleay, 1871 [species]

Onthophagus incostatus D'Orbigny, 1902 [species]

Onthophagus indigus Péringuey, 1901 [species]

Onthophagus indosinicus Boucomont, 1921 [species]

Onthophagus indutus D'orbigny, 1902 [species]

Onthophagus inediapterus Kohlmann & Solis, 2001 [species]

Onthophagus inelegans Balthasar, 1935 [species]

Onthophagus ineptus Harold, 1871 [species]

Onthophagus inermiceps D'orbigny, 1905 [species]

Onthophagus inermicollis D'orbigny, 1908 [species]

Onthophagus inermivertex Boucomont, 1921 [species]

Onthophagus infaustus Cambefort, 1984 [species]

Onthophagus inflaticollis Bates, 1887 [species]

Onthophagus inflatus WALTER & CAMBEFORT, 1977 [species]

Onthophagus infucatus Harold, 1877 [species]

Onthophagus infuscatus Klug, 1845 [species]

Onthophagus insignicollis Frey, 1954 [species]

Onthophagus insignis Péringuey, 1896 [species]

Onthophagus insularis Boheman, 1858 [species]

Onthophagus insulindicus Boucomont, 1914 [species]

Onthophagus insulsus Péringuey, 1901 [species]

Onthophagus intermixtus D'orbigny, 1902 [species]

Onthophagus interruptus Raffray, 1877 [species]

Onthophagus interstitialis Fahraeus, 1857 [species]

Onthophagus intonsus D'orbigny, 1913 [species]

Onthophagus intricatus Moretto, 2010 [species]

Onthophagus investigator Lansberge, 1885 [species]

Onthophagus investis D'orbigny, 1904 [species]

Onthophagus iodiellus Bates, 1887 [species]

Onthophagus irianus Balthasar, 1969 [species]

Onthophagus iris Sharp, 1875 [species]

Onthophagus isanus Masumoto, Ochi & Hanboonsong, 2008 [species]

Onthophagus ishiii OCHI & KON, 1995 [species]

Onthophagus isikdagensis Pittino, 2004 [species]

Onthophagus itiokai Ochi, Kon & Kishimoto-Yamada, 2010 [species]

Onthophagus itoi Nomura, 1976 [species]

Onthophagus iulicola Cambefort, 1984 [species]

Onthophagus iumienus Masumoto, Ochi & Hanboonsong, 2007 [species]

Onthophagus iyengari Arrow, 1931 [species]

Onthophagus jacksoni D'orbigny, 1913 [species]

Onthophagus jacobeus Boucomont, 1924 [species]

Onthophagus jalamari Matthews, 1972 [species]

Onthophagus jalapensis Balthasar, 1939 [species]

Onthophagus jangga Matthews, 1972 [species]

Onthophagus janssensi Frey, 1958 [species]

Onthophagus janthinus Harold, 1875 [species]

Onthophagus janushevi Kabakov, 1983 [species]

Onthophagus japonicus Harold, 1875 [species]

Onthophagus javacupreus Krikken & Huijbregts, 2009 [species]

Onthophagus javaecola Balthasar, 1959 [species]

Onthophagus javanensis Balthasar, 1963 [species]

Onthophagus javanitidus Ochi, Kon & Hartini, 2010 [species]

Onthophagus javanoides Huijbregts & Krikken, 2011 [species]

Onthophagus jeanneli D'orbigny, 1913 [species]

Onthophagus jeannelianus Paulian, 1945 [species]

Onthophagus jingping Masumoto, Ochi & Hanboonsong, 2007 [species]

Onthophagus joannae Goljan, 1953 [species]

Onthophagus johkii Ochi & Kon, 1994 [species]

Onthophagus joliveti Paulian, 1972 [species]

Onthophagus jubatus Harold, 1869 [species]

Onthophagus jugicola D'orbigny, 1902 [species]

Onthophagus juncticornis D'orbigny, 1908 [species]

Onthophagus justei Walter, 1989 [species]

Onthophagus juvencus Klug, 1835 [species]

Onthophagus kabakovi Martin-Piera, 1985 [species]

Onthophagus kabulicus Balthasar, 1971 [species]

Onthophagus kachinicus Kabakov, 2006 [species]

Onthophagus kachowskii Olsoufief, 1900 [species]

Onthophagus kaengkrachangus Masumoto, Ochi & Hanboonsong, 2008 [species]

Onthophagus kakadu STOREY & WEIR, 1990 [species]

Onthophagus kanarensis Arrow, 1931 [species]

Onthophagus kangeanus Paulian, 1936 [species]

Onthophagus kanyaayonus Masumoto, 1992 [species]

Onthophagus kaosoidowensis Masumoto, Ochi & Hanboonsong, 2008 [species]

Onthophagus kapitensis Frey, 1971 [species]

Onthophagus kapuri Endrödi, 1974 [species]

Onthophagus karenensis Masumoto, Ochi & Hanboonsong, 2008 [species]

Onthophagus kashizakii Kon & Ochi, 2005 [species]

Onthophagus kashmirensis Balthasar, 1966 [species]

Onthophagus kassaicus D'orbigny, 1908 [species]

Onthophagus katangensis Frey, 1961 [species]

Onthophagus katoi Ochi & Araya, 1996 [species]

Onthophagus katualensis Quedenfeldt, 1888 [species]

Onthophagus kavirondus D'orbigny, 1915 [species]

Onthophagus kawaharai Ochi & Kon, 2007 [species]

Onthophagus kchatriya Boucomont, 1914 [species]

Onthophagus keiseri Frey, 1956 [species]

Onthophagus kentingensis Nomura, 1973 [species]

Onthophagus keralensis Frey, 1975 [species]

Onthophagus keralicus Biswas & Chatterjee, 1986 [species]

Onthophagus khonkaenus Masumoto, Ochi & Hanboonsong, 2008 [species]

Onthophagus khonmiinitnoi Masumoto, 1990 [species]

Onthophagus kiambram Storey, 1977 [species]

Onthophagus kikutai Ochi & Kon, 2005 [species]

Onthophagus kilimanus Kolbe, 1891 [species]

Onthophagus kimioi Ochi, Kon & Kawahara, 2011 [species]

Onthophagus kindermanni Harold, 1877 [species]

Onthophagus kindianus Frey, 1953 [species]

Onthophagus kingstoni Cambefort, 1980 [species]

Onthophagus kinhthaicus Kabakov, 1998 [species]

Onthophagus kirki Frey, 1975 [species]

Onthophagus kirokanus Frey, 1975 [species]

Onthophagus kiuchianus Masumoto, Yang & Ochi, 2004 [species]

Onthophagus kiuchii Masumoto, 1995 [species]

Onthophagus kiyoshii Ochi & Kon, 2007 [species]

Onthophagus klapperichi Balthasar, 1952 [species]

Onthophagus kleinei Balthasar, 1935 [species]

Onthophagus knapperti Krikken, 1981 [species]

Onthophagus kochi Frey, 1957 [species]

Onthophagus koebelei Blackburn, 1903 [species]

Onthophagus koechlei Masumoto, Ochi & Kon, 2011 [species]

Onthophagus koichii Ochi & Araya, 1992 [species]

Onthophagus kokereka Matthews, 1972 [species]

Onthophagus kolaka Huijbregts & Krikken, 2009 [species]

Onthophagus kolbei Marcus, 1921 [species]

Onthophagus kolenatii Reitter, 1893 [species]

Onthophagus koma Matsumura, 1937 [species]

Onthophagus komareki Balthasar, 1935 [species]

Onthophagus kondaoensis Kabakov, 1994 [species]

Onthophagus koni Ochi, 2007 [species]

Onthophagus konoi Matsumura, 1938 [species]

Onthophagus konsarnensis Masumoto, Ochi & Hanboonsong, 2008 [species]

Onthophagus kontumicus Kabakov, 1983 [species]

Onthophagus kora Storey, 1977 [species]

Onthophagus koryoensis Kim, 1985 [species]

Onthophagus koshunensis Balthasar, 1941 [species]

Onthophagus kouassii Cambefort, 1984 [species]

Onthophagus kozlovi Kabakov, 1990 [species]

Onthophagus kraatzeanus Lansberge, 1883 [species]

Onthophagus krakadaakhomus Masumoto, 1992 [species]

Onthophagus kryzhanovskii Kabakov, 1982 [species]

Onthophagus kuatunensis Balthasar, 1942 [species]

Onthophagus kukunorensis Kabakov, 1990 [species]

Onthophagus kulti Balthasar, 1952 [species]

Onthophagus kuluensis Bates, 1891 [species]

Onthophagus kulzeri Frey, 1958 [species]

Onthophagus kumaonensis Arrow, 1931 [species]

Onthophagus kumbaingeri Matthews, 1972 [species]

Onthophagus kwaleensis Moretto & Nicolas, 2002 [species]

Onthophagus kyleensis Frey, 1975 [species]

Onthophagus labdacus Balthasar, 1969 [species]

Onthophagus laborans Arrow, 1931 [species]

Onthophagus laceratus Gerstaecker, 1871 [species]

Onthophagus lacustris Harold, 1877 [species]

Onthophagus laetus D'orbigny, 1913 [species]

Onthophagus laevatus D'orbigny, 1902 [species]

Onthophagus laeviceps D'orbigny, 1902 [species]

Onthophagus laevicollis Lansberge, 1883 [species]

Onthophagus laevidorsis D'orbigny, 1907 [species]

Onthophagus laevigatus Fabricius, 1798 [species]

Onthophagus laevis Harold, 1880 [species]

O. l. asiaticus, O. l. lampromelas, O. l. stevensi

Onthophagus laevissimus D'orbigny, 1905 [species]

Onthophagus lagnyi Paulian, 1937 [species]

Onthophagus lahorensis Kabakov, 2008 [species]

Onthophagus lakyim Masumoto, 1990 [species]

Onthophagus lallieri d'Orbigny, 1902 [species]

Onthophagus lallierianus Janssens, 1953 [species]

Onthophagus lamellicornis Frey, 1953 [species]

Onthophagus lamelliger Gerstaecker, 1871 [species]

Onthophagus lamgalio Matthews, 1972 [species]

Onthophagus laminatus Macleay, 1864 [species]

Onthophagus laminicornis Lansberge, 1886 [species]

Onthophagus laminidorsis D'orbigny, 1902 [species]

Onthophagus laminosus D'orbigny, 1905 [species]

Onthophagus lamnifer D'orbigny, 1902 [species]

Onthophagus lamotellus Cambefort, 1980 [species]

Onthophagus lamottei Frey, 1962 [species]

Onthophagus lamtoensis Cambefort, 1984 [species]

Onthophagus lamtoi Cambefort, 1980 [species]

Onthophagus lamyi Paulian, 1945 [species]

Onthophagus lanista Castelnau, 1840 [species]

Onthophagus lannamiibun Masumoto, Ochi & Hanboonsong, 2002 [species]

Onthophagus laotianus Boucomont, 1919 [species]

Onthophagus lapillus Arrow, 1931 [species]

Onthophagus laratinus Arrow, 1916 [species]

Onthophagus lassulus Balthasar, 1964 [species]

Onthophagus latenasutus Arrow, 1941 [species]

Onthophagus latepunctatus D'orbigny, 1913 [species]

Onthophagus latestriatus D'orbigny, 1908 [species]

Onthophagus latevittatus D'orbigny, 1905 [species]

Onthophagus laticollis Klug, 1835 [species]

Onthophagus laticornis Gebler, 1823 [species]

Onthophagus latigena D'Orbigny, 1897 [species]

Onthophagus latigibber D ORBIGNY, 1902 [species]

Onthophagus latipennis D'orbigny, 1897 [species]

Onthophagus latissimus D'orbigny, 1902 [species]

Onthophagus latro Harold, 1877 [species]

Onthophagus leai Blackburn, 1895 [species]

Onthophagus leanus Goidanich, 1926 [species]

Onthophagus lebasi Boucomont, 1932 [species]

Onthophagus lefebvrei D'orbigny, 1913 [species]

Onthophagus lefiniensis Balthasar, 1967 [species]

Onthophagus legendrei WALTER & CAMBEFORT, 1977 [species]

Onthophagus lemagneni D'orbigny, 1902 [species]

Onthophagus lemekensis D'orbigny, 1915 [species]

Onthophagus lemniscatus Gillet, 1924 [species]

Onthophagus lemur Fabricius 1781 [species]

Onthophagus lemuroides D'orbigny, 1898 [species]

Onthophagus lenis Kabakov, 1998 [species]

Onthophagus lenzi Harold, 1875 [species]

Onthophagus leomontanus Boucomont, 1914 [species]

Onthophagus leroyi D'orbigny, 1902 [species]

Onthophagus leucopygus Harold, 1867 [species]

Onthophagus leucostigma Steven, 1811 [species]

Onthophagus leusermontis Huijbregts & Krikken, 2011 [species]

Onthophagus leytensis Ochi & Araya, 1992 [species]

Onthophagus liberianus Lansberge, 1883 [species]

Onthophagus liewi Ochi & Kon, 2005 [species]

Onthophagus lilliputanus Lansberge, 1883 [species]

Onthophagus limbatus Herbst, 1789 [species]

Onthophagus limonensis Kohlmann & Solis, 2001 [species]

Onthophagus lindaae Masumoto, 1989 [species]

Onthophagus lindu Huijbregts & Krikken, 2009 [species]

Onthophagus liodermus D'orbigny, 1913 [species]

Onthophagus lioides D'orbigny, 1902 [species]

Onthophagus liopterus Harold, 1880 [species]

Onthophagus liothorax Koshantschikov, 1894 [species]

Onthophagus lituratus Roth, 1851 [species]

Onthophagus liwagensis Ochi & Kon, 2006 [species]

Onthophagus lobi Cambefort, 1984 [species]

Onthophagus lojanus Balthasar, 1939 [species]

Onthophagus lomii Müller, 1942 [species]

Onthophagus longefossus D'Orbigny, 1905 [species]

Onthophagus longegranus Frey, 1957 [species]

Onthophagus longiceps D'orbigny, 1904 [species]

Onthophagus longimanus Bates, 1887 [species]

Onthophagus longipes Paulian, 1937 [species]

Onthophagus longipilis D'orbigny, 1905 [species]

Onthophagus lore Huijbregts & Krikken, 2009 [species]

Onthophagus lorianus Gillet, 1930 [species]

Onthophagus loricatus Klug, 1855 [species]

Onthophagus loroi Balthasar, 1941 [species]

Onthophagus loudetiae Cambefort, 1984 [species]

Onthophagus loveni Gillet, 1928 [species]

Onthophagus loxodontae Cambefort, 1984 [species]

Onthophagus loxodontaphilus Moretto, 2009 [species]

Onthophagus lucasi D'Orbigny, 1902 [species]

Onthophagus lucidus Illiger, 1800 [species]

Onthophagus luctuosus Boucomont, 1914 [species]

Onthophagus ludicrus Balthasar, 1969 [species]

Onthophagus ludio Boucomont, 1914 [species]

Onthophagus lugens Fahraeus, 1857 [species]

Onthophagus lugubris Fahraeus, 1857 [species]

Onthophagus luismargaritorum Delgado, 1995 [species]

Onthophagus lujendae Bates, 1888 [species]

Onthophagus lumareti WALTER & CAMBEFORT, 1977 [species]

Onthophagus lunatus Harold, 1868 [species]

Onthophagus lunulifer Boucomont, 1914 [species]

Onthophagus luridipennis Boheman, 1858 [species]

Onthophagus lutaticollis D'orbigny, 1907 [species]

Onthophagus luteosignatus Lansberge, 1883 [species]

Onthophagus lutosopictus Fairmaire, 1897 [species]

Onthophagus macedonicus Miksic, 1959 [species]

Onthophagus macleayi Blackburn, 1903 [species]

Onthophagus macrocephalus Kirby, 1818 [species]

Onthophagus macroliberianus Moretto, 2010 [species]

Onthophagus macrothorax D'orbigny, 1902 [species]

Onthophagus maculatus Fabricius, 1801 [species]

Onthophagus maculosipennis Gillet, 1930 [species]

Onthophagus maculosus D'orbigny, 1908 [species]

Onthophagus madoqua Arrow, 1931 [species]

Onthophagus maephaluangus Masumoto, Ochi & Hanboonsong, 2007 [species]

Onthophagus maesaensis Masumoto, Ochi & Hanboonsong, 2002 [species]

Onthophagus magnini Paulian, 1933 [species]

Onthophagus magnioculus Ochi & Kon, 2006 [species]

Onthophagus magnipygus Boucomont, 1914 [species]

Onthophagus mairuu Masumoto, 1989 [species]

Onthophagus maki Illiger, 1803 [species]

Onthophagus makokou Walter, 1989 [species]

Onthophagus malabarensis Boucomont, 1919 [species]

Onthophagus malangensis Boucomont, 1914 [species]

Onthophagus malasiacus Gillet, 1927 [species]

Onthophagus maleengnaafon Masumoto, 1990 [species]

Onthophagus maleengnoi Masumoto, 1990 [species]

Onthophagus malevolus Balthasar, 1964 [species]

Onthophagus malthinus Gillet, 1930 [species]

Onthophagus mamillatus Lea, 1923 [species]

Onthophagus manguliensis Boucomont, 1914 [species]

Onthophagus manipurensis Arrow, 1907 [species]

Onthophagus maniti Masumoto, 1989 [species]

Onthophagus mankonoensis Balthasar, 1966 [species]

Onthophagus manya Matthews, 1972 [species]

Onthophagus marahouensis Cambefort, 1984 [species]

Onthophagus margaretensis Blackburn, 1903 [species]

Onthophagus margaritifer D'orbigny, 1898 [species]

Onthophagus marginalis Gebler, 1817 [species]

O. m. andalusicus, O. m. marginalis, O. m. przewalski

Onthophagus marginatus Castelnau, 1840 [species]

Onthophagus marginicollis Harold, 1880 [species]

Onthophagus marginidens D'Orbigny, 1904 [species]

Onthophagus marginifer Frey, 1953 [species]

Onthophagus maringouensis Palestrini, 1992 [species]

Onthophagus mariozuninoi Delgado, Navarrete & Blackaller-Bages, 1993 [species]

Onthophagus marmotae Kabakov, 1990 [species]

Onthophagus marshalli D'orbigny, 1908 [species]

Onthophagus martelli Frey, 1972 [species]

Onthophagus martialis Boucomont, 1914 [species]

Onthophagus maryatiae Ochi & Kon, 2005 [species]

Onthophagus masahiroi Masumoto, Ochi & Hanboonsong, 2008 [species]

Onthophagus masaicus D'orbigny, 1905 [species]

Onthophagus masaoi Ochi, 1992 [species]

Onthophagus massai Baraud, 1975 [species]

Onthophagus masumotoi Ochi, 1985 [species]

Onthophagus matae Cambefort, 1984 [species]

Onthophagus matanyo Huijbregts & Krikken, 2009 [species]

Onthophagus matsudai Ochi & Kon, 2004 [species]

Onthophagus matsuii Ochi & Kon, 2006 [species]

Onthophagus mauritii Boucomont, 1919 [species]

Onthophagus maya Zunino, 1981 [species]

Onthophagus mayeri Harold, 1876 [species]

Onthophagus mediofuscatus D'Orbigny, 1908 [species]

Onthophagus medius Kugelann, 1792 [species]

Onthophagus medvedevi Kabakov, 1982 [species]

Onthophagus megapacificus Ochi & Kon, 2006 [species]

Onthophagus megathorax Gillet, 1921 [species]

Onthophagus mekara Huijbregts & Krikken, 2009 [species]

Onthophagus melanocephalus Klug, 1845 [species]

Onthophagus melitaeus Fabricius 1798 [species]

Onthophagus mendeli Ochi, Kon & Barclay, 2009 [species]

Onthophagus mendicus Gillet, 1924 [species]

Onthophagus menieri Ochi, 2003 [species]

Onthophagus menkaoensis WALTER & CAMBEFORT, 1977 [species]

Onthophagus mentaveiensis Boucomont, 1914 [species]

Onthophagus merdarius Chevrolat, 1865 [species]

Onthophagus merdrignaci WALTER & CAMBEFORT, 1977 [species]

Onthophagus meruanus D'orbigny, 1908 [species]

Onthophagus merus Péringuey, 1901 [species]

Onthophagus metalliceps Arrow 1931 [species]

Onthophagus metalliger D'orbigny, 1913 [species]

Onthophagus mexicanus Bates, 1887 [species]

Onthophagus micropterus ZUNINO & HALFFTER, 1981 [species]

Onthophagus micros D'orbigny, 1902 [species]

Onthophagus mije STOREY & WEIR, 1990 [species]

Onthophagus miles D'orbigny, 1905 [species]

Onthophagus militaris Boucomont, 1914 [species]

Onthophagus millamilla Matthews, 1972 [species]

Onthophagus millingeni D'orbigny, 1898 [species]

Onthophagus mimikanus Balthasar, 1969 [species]

Onthophagus mimus D'orbigny, 1913 [species]

Onthophagus minans D'orbigny, 1913 [species]

Onthophagus minax D'orbigny, 1904 [species]

Onthophagus mindanaensis Boucomont, 1914 [species]

Onthophagus mindanaoensis Frey, 1971 [species]

Onthophagus minettii Moretto & Nicolas, 2002 [species]

Onthophagus minotaurus Arrow, 1941 [species]

Onthophagus minutissimus D'orbigny, 1908 [species]

Onthophagus minutulus Harold, 1875 [species]

Onthophagus minutus Hausmann, 1807 [species]

Onthophagus mirabilis Bates, 1887 [species]

Onthophagus mirandus Arrow, 1931 [species]

Onthophagus miricollis Frey, 1962 [species]

Onthophagus miricornis d'Orbigny, 1902 [species]

Onthophagus mirifrons D'orbigny, 1905 [species]

Onthophagus miscellaneus D'orbigny, 1908 [species]

Onthophagus miscellus D'orbigny, 1905 [species]

Onthophagus misellus D'orbigny, 1907 [species]

Onthophagus missor Balthasar, 1938 [species]

Onthophagus mixticeps D'orbigny, 1905 [species]

Onthophagus mixtidorsis D'orbigny, 1905 [species]

Onthophagus mixtifrons D'Orbigny, 1910 [species]

Onthophagus miyakei Ochi & Araya, 1992 [species]

Onthophagus mjobergi Gillet, 1925 [species]

Onthophagus mniszechi Harold, 1869 [species]

Onthophagus moajat Huijbregts & Krikken, 2009 [species]

Onthophagus mocquerysi D'orbigny, 1902 [species]

Onthophagus modestus Harold, 1862 [species]

Onthophagus mogo Krikken & Huijbregts, 2008 [species]

Onthophagus monardi Boucomont, 1936 [species]

Onthophagus monardiellus Frey, 1963 [species]

Onthophagus monforti Josso & Prévost, 2006 [species]

Onthophagus mongana STOREY & WEIR, 1990 [species]

Onthophagus mongkhoni Masumoto, Hanboonsong & Ochi, 2002 [species]

Onthophagus monochromus D'Orbigny, 1913 [species]

Onthophagus monodon Fahraeus, 1857 [species]

Onthophagus monteithi Matthews, 1972 [species]

Onthophagus monticupreus Krikken & Huijbregts, 2009 [species]

Onthophagus montishannoniae Krikken & Huijbrechts, 2008 [species]

Onthophagus montivagus D'orbigny, 1902 [species]

Onthophagus montreuili Moretto & Génier, 2010 [species]

Onthophagus mopsus Fabricius, 1792 [species]

Onthophagus mordindangus Masumoto, Ochi & Hanboonsong, 2008 [species]

Onthophagus moreleti Baraud, 1980 [species]

Onthophagus moroni ZUNINO & HALFFTER, 1988 [species]

Onthophagus morosus Gerstaecker, 1871 [species]

Onthophagus mouhoti Harold, 1875 [species]

Onthophagus mpassa Walter, 1982 [species]

Onthophagus mucronatus Thomson, 1858 [species]

Onthophagus mucronifer D'orbigny, 1905 [species]

Onthophagus muelleri Lansberge, 1883 [species]

Onthophagus mulgravei Paulian, 1937 [species]

Onthophagus multicornis D'Orbigny, 1908 [species]

Onthophagus multipunctatus Boucomont, 1914 [species]

Onthophagus mundill Matthews, 1972 [species]

Onthophagus murasakianus Nomura, 1976 [species]

O. m. carnarius, O. m. miyakoinsularis

Onthophagus murchisoni Blackburn, 1892 [species]

Onthophagus musculus Frey, 1963 [species]

O. m. ganganensis

Onthophagus mushensis Matsumura, 1938 [species]

Onthophagus mutatus Harold, 1859 [species]

Onthophagus muticifrons Endrödi, 1973 [species]

Onthophagus muticus Macleay, 1864 [species]

Onthophagus naaroon Masumoto, 1990 [species]

Onthophagus nabeleki Balthasar, 1939 [species]

Onthophagus naevius D'orbigny, 1913 [species]

Onthophagus naevuliger D'orbigny, 1908 [species]

Onthophagus nagasawai Matsumura, 1938 [species]

Onthophagus nagpurensis Arrow, 1931 [species]

Onthophagus nammuldi Matthews, 1972 [species]

Onthophagus namnaoensis Masumoto & Ochi & Hanboonsong, 2002 [species]

Onthophagus nampatensis Tarasov & Kabakov, 2010 [species]

Onthophagus nanus Harold, 1878 [species]

Onthophagus napolovi Kabakov, 1998 [species]

Onthophagus nasalis Arrow, 1931 [species]

Onthophagus nasicornis Harold, 1869 [species]

Onthophagus nasicus Gillet, 1930 [species]

Onthophagus nasidens D'Orbigny, 1905 [species]

Onthophagus naso Fahraeus, 1857 [species]

Onthophagus nasonis Kabakov, 1998 [species]

Onthophagus nasutus Guérin-Méneville, 1855 [species]

Onthophagus natronis Cambefort, 1984 [species]

Onthophagus navarretorum DELGADO & CAPISTAN, 1996 [species]

Onthophagus naviculifer Boucomont, 1919 [species]

Onthophagus neboissi Frey, 1970 [species]

Onthophagus nebulosus Reiche, 1864 [species]

Onthophagus necessarius Reitter, 1893 [species]

Onthophagus necrophagus Arrow, 1931 [species]

Onthophagus neervoorti Boucomont, 1914 [species]

Onthophagus nefarius Balthasar, 1941 [species]

Onthophagus negligens Walker, 1858 [species]

Onthophagus negrosensis Ochi & Araya, 1992 [species]

Onthophagus negus Raffray, 1882 [species]

Onthophagus nemorivagus Kohlmann & Solis, 2001 [species]

Onthophagus neocolobus Scheuern, 1996 [species]

Onthophagus neofurcatus Goidanich, 1926 [species]

Onthophagus neomirabilis Howden, 1973 [species]

Onthophagus neostenocerus Goidanich, 1926 [species]

Onthophagus ngottoensis Moretto & Nicolas, 2002 [species]

Onthophagus niasensis Boucomont, 1914 [species]

Onthophagus nicobaricus Biswas, Chatterjee & Sengupta, 1999 [species]

Onthophagus nicolasi Moretto, 2004 [species]

Onthophagus nigellus Illiger, 1803 [species]

Onthophagus nigerianus D'orbigny, 1913 [species]

Onthophagus nigriceps Raffray, 1877 [species]

Onthophagus nigricornis Fairmaire, 1887 [species]

Onthophagus nigrinus Paulian, 1937 [species]

Onthophagus nigriobscurior Ochi, Kon & Tsubaki, 2009 [species]

Onthophagus nigripennis D'orbigny, 1908 [species]

Onthophagus nigriventris D'orbigny, 1905 [species]

Onthophagus nigrivestis D'Orbigny, 1902 [species]

Onthophagus nigropubens D'orbigny, 1908 [species]

Onthophagus nigroviolaceus d'Orbigny 1902 [species]

Onthophagus nikolajevi Kabakov, 2006 [species]

Onthophagus nilgirensis Gillet, 1922 [species]

Onthophagus nilicola D'orbigny, 1902 [species]

Onthophagus niloticus Harold, 1879 [species]

Onthophagus nimbatus D'orbigny, 1905 [species]

Onthophagus niokolokoba Moretto, 2007 [species]

Onthophagus nitefactus Harold, 1877 [species]

Onthophagus nitidifrons D'orbigny, 1905 [species]

Onthophagus nitidior Bates, 1887 [species]

Onthophagus nitidulus Klug, 1845 [species]

Onthophagus nitidus Waterhouse, 1875 [species]

Onthophagus nodieri D'orbigny, 1908 [species]

Onthophagus nodulifer Harold, 1867 [species]

Onthophagus nongkaiensis Masumoto, Ochi & Hanboonsong, 2002 [species]

Onthophagus nonstriatus D'orbigny, 1905 [species]

Onthophagus notatus D'orbigny, 1902 [species]

Onthophagus notiodes Solis & Kohlmann, 2003 [species]

Onthophagus novaeirlandiae Balthasar, 1969 [species]

Onthophagus nuba D'Orbigny, 1902 [species]

Onthophagus nubilus Kohlmann & Solis, 2001 [species]

Onthophagus nudatus D'orbigny, 1913 [species]

Onthophagus nudifrons Balthasar, 1939 [species]

Onthophagus nudus D'orbigny, 1908 [species]

Onthophagus numidicus d'Orbigny 1908 [species]

Onthophagus nurestanicus Kabakov, 1982 [species]

Onthophagus nurubuan Matthews, 1972 [species]

Onthophagus nusatenggaricus Krikken & Huijbreghts, 2008 [species]

Onthophagus nyctopus Bates, 1887 [species]

Onthophagus nymani Gillet, 1930 [species]

Onthophagus obenbergeri Balthasar, 1965 [species]

Onthophagus oberthuri D'orbigny, 1898 [species]

Onthophagus obliquus Olivier, 1789 [species]

Onthophagus obliviosus Balthasar, 1935 [species]

Onthophagus oblongus Zhang, 1997 [species]

Onthophagus obscurior Boucomont, 1914 [species]

Onthophagus obtusicornis Fahraeus, 1857 [species]

Onthophagus obtutus Péringuey, 1901 [species]

Onthophagus occipitalis Lansberge, 1885 [species]

Onthophagus ocellatopunctatus Waterhouse, 1875 [species]

Onthophagus ocellidorsis D'orbigny, 1915 [species]

Onthophagus ocellifer D'orbigny, 1902 [species]

Onthophagus ocelliger Harold, 1877 [species]

Onthophagus ochii Masumoto, 1988 [species]

Onthophagus ochreatus D'Orbigny, 1897 [species]

Onthophagus ochromerus Harold, 1877 [species]

Onthophagus ochropygus D'orbigny, 1902 [species]

Onthophagus octogonus Frey, 1962 [species]

Onthophagus octonaevus Kabakov, 1994 [species]

Onthophagus oculatus Arrow 1931 [species]

Onthophagus ohbayashii Nomura, 1939 [species]

Onthophagus ohkuboi Ochi & Kon, 2006 [species]

Onthophagus ohmomoi Miyake, 2000 [species]

Onthophagus okahandjanus Balthasar, 1974 [species]

Onthophagus olidus Balthasar, 1964 [species]

Onthophagus olsoufieffi Boucomont, 1924 [species]

Onthophagus omostigma D'orbigny, 1902 [species]

Onthophagus onorei ZUNINO & HALFFTER, 1997 [species]

Onthophagus onthochromus Arrow, 1913 [species]

Onthophagus ontosatu Huijbregts & Krikken, 2012 [species]

Onthophagus opacicollis Reitter, 1893 [species]

Onthophagus opacifalculatus Ochi, Kon & Tsubaki, 2009 [species]

Onthophagus ophion Erichson, 1847 [species]

O. o. confusus

Onthophagus ophtalmicus Frey, 1958 [species]

Onthophagus opulentus D'Orbigny, 1902 [species]

Onthophagus orbicularis Lansberge, 1885 [species]

Onthophagus orbus Boucomont, 1924 [species]

Onthophagus orientalis Harold, 1868 [species]

Onthophagus orissanus Arrow, 1931 [species]

Onthophagus ornaticollis Gillet, 1930 [species]

Onthophagus ornatulus D'orbigny, 1908 [species]

Onthophagus orphnoides Bates, 1887 [species]

Onthophagus orthocerus Thomson, 1858 [species]

Onthophagus osculatii Guérin-Méneville, 1855 [species]

Onthophagus osellai Pittino, 1982 [species]

Onthophagus oshimanus Nakane, 1960 [species]

Onthophagus otai Ochi & Kon, 2006 [species]

Onthophagus otjivarongus Balthasar, 1967 [species]

Onthophagus ouratita Matthews, 1972 [species]

Onthophagus ovatus Linnaeus 1767 [species]

Onthophagus overlaeti Janssens, 1953 [species]

Onthophagus ovigranosus Frey, 1971 [species]

Onthophagus ovulum Gerstaecker, 1871 [species]

Onthophagus pacificus Lansberge, 1885 [species]

Onthophagus pactolus Fabricius, 1787 [species]

Onthophagus padrinoi Delgado, 1999 [species]

Onthophagus palatus Boucomont, 1914 [species]

Onthophagus palavanus Balthasar, 1959 [species]

Onthophagus palawanicus Ochi & Kon, 2004 [species]

Onthophagus palestriniae Moretto & Nicolas, 2004 [species]

Onthophagus paliceps Arrow, 1931 [species]

Onthophagus pallens D'orbigny, 1908 [species]

Onthophagus pallidipennis Fahraeus, 1857 [species]

Onthophagus pallidus D'orbigny, 1905 [species]

Onthophagus paluma Matthews, 1972 [species]

Onthophagus panfilovi Kabakov, 1983 [species]

Onthophagus panici Petrovitz, 1964 [species]

Onthophagus panoplus Bates, 1888 [species]

Onthophagus paolae Palestrini, 1992 [species]

Onthophagus papuensis Harold, 1877 [species]

Onthophagus papulatorius Kabakov, 2008 [species]

Onthophagus papulatus Boucomont, 1914 [species]

Onthophagus parachandrai Ochi, Kon & Tsubaki, 2009 [species]

Onthophagus parafalculatus Ochi, Kon & Tsubaki, 2009 [species]

Onthophagus parafasciatus Balthasar, 1974 [species]

Onthophagus parallelicornis Macleay, 1887 [species]

Onthophagus paramasaoi Ochi, Kon & Barclay, 2009 [species]

Onthophagus parapalatus KRIKKEN & HUIJBREGTS, 1988 [species]

Onthophagus parapedisequus Balthasar, 1969 [species]

Onthophagus paravinctus Tagliaferri & Moretto, 2012 [species]

Onthophagus parceguttatus D'orbigny, 1902 [species]

Onthophagus parcenotatus D'orbigny, 1913 [species]

Onthophagus parcepictus D'orbigny, 1908 [species]

Onthophagus parcepilosus Balthasar, 1966 [species]

Onthophagus pardalis Fabricius 1798 [species]

Onthophagus parenthesis Boucomont, 1913 [species]

Onthophagus parisii Balthasar, 1941 [species]

Onthophagus parmatus Reitter, 1892 [species]

Onthophagus paroculus Krikken & Huijbregts, 1987 [species]

Onthophagus parrumbal Matthews, 1972 [species]

Onthophagus parryi Harold, 1869 [species]

Onthophagus parumnotatus Fahraeus, 1857 [species]

Onthophagus parvidens Frey, 1971 [species]

Onthophagus parvifrons D'orbigny, 1907 [species]

Onthophagus parviobscurior Ochi, Kon & Tsubaki, 2009 [species]

Onthophagus parvulus Fabricius, 1798 [species]

Onthophagus parvus Blanchard, 1853 [species]

Onthophagus pastillatus Boucomont, 1919 [species]

Onthophagus patinatus Marcus, 1917 [species]

Onthophagus paucigranosus Lansberge, 1886 [species]

Onthophagus paulianellus Frey, 1958 [species]

Onthophagus pauliani Balthasar, 1937 [species]

Onthophagus pauper Boucomont, 1914 [species]

Onthophagus pauxillus D'orbigny, 1902 [species]

Onthophagus pavidus Harold, 1877 [species]

Onthophagus pedator Sharp, 1875 [species]

Onthophagus pedester Howden & Génier, 2004 [species]

Onthophagus pedisequus Balthasar, 1969 [species]

Onthophagus pegesimallus Marcus, 1917 [species]

Onthophagus pendjarius Josso & Prévost, 2006 [species]

Onthophagus penicillatus Harold, 1879 [species]

Onthophagus peninsularis Boucomont, 1914 [species]

Onthophagus peninsulocupreus Krikken & Huijbregts, 2009 [species]

Onthophagus peninsulomerus Huijbregts & Krikken, 2011 [species]

Onthophagus penmani Masumoto & Ochi & Hanboonsong, 2002 [species]

Onthophagus pentacanthus Harold, 1867 [species]

Onthophagus peramelinus Lea, 1923 [species]

Onthophagus perniger Boucomont, 1930 [species]

Onthophagus perpilosus Macleay, 1871 [species]

Onthophagus personatus Boucomont, 1914 [species]

Onthophagus petenensis HOWDEN & GILL, 1993 [species]

Onthophagus petrovitzi Frey, 1975 [species]

Onthophagus petrovitzianus Pittino, 2006 [species]

Onthophagus pexatus Harold, 1869 [species]

Onthophagus peyrierasi Paulian, 1985 [species]

Onthophagus phanaeicollis Lansberge, 1883 [species]

Onthophagus phanaeides Frey, 1956 [species]

Onthophagus phanaeiformis Boucomont, 1914 [species]

Onthophagus phanaeomorphus Janssens, 1954 [species]

Onthophagus phatoensis Masumoto, Ochi & Hanboonsong, 2008 [species]

Onthophagus phetchabunensis Masumoto, Ochi & Hanboonsong, 2002 [species]

Onthophagus philippinensis Boucomont, 1919 [species]

Onthophagus phillippsorum Krikken & Huijbregts, 1987 [species]

Onthophagus phoenicocerus Lea, 1923 [species]

Onthophagus phraoensis Masumoto, 1990 [species]

Onthophagus phrixus Balthasar, 1969 [species]

Onthophagus phrutsaphaakhomus Masumoto, 1992 [species]

Onthophagus phukhieoensis Masumoto, Ochi & Hanboonsong, 2007 [species]

Onthophagus phuquoci Paulian, 1945 [species]

Onthophagus picatus D'orbigny, 1902 [species]

Onthophagus piceiceps D'orbigny, 1904 [species]

Onthophagus piceorufulus Kabakov, 1994 [species]

Onthophagus picipennis Hope, 1841 [species]

Onthophagus pictipennis D'orbigny, 1913 [species]

Onthophagus pictipodex D'orbigny, 1902 [species]

Onthophagus picturatus D'orbigny, 1908 [species]

Onthophagus pictus Reitter, 1893 [species]

Onthophagus piffli Petrovitz, 1961 [species]

Onthophagus pilicollis D'orbigny, 1902 [species]

Onthophagus pilipodex D'orbigny, 1913 [species]

Onthophagus pillara Matthews, 1972 [species]

Onthophagus pilosus Fahraeus, 1857 [species]

Onthophagus pilularius Lansberge, 1883 [species]

Onthophagus pimpasaleei Masumoto, Ochi & Hanboonsong, 2008 [species]

Onthophagus pinaroo STOREY & WEIR, 1990 [species]

Onthophagus pinguis Gerstaecker, 1871 [species]

Onthophagus pipitzi Ancey, 1883 [species]

Onthophagus pisciphagus WALTER & CAMBEFORT, 1977 [species]

Onthophagus piyawati Masumoto, Ochi & Hanboonsong, 2002 [species]

Onthophagus placens Péringuey, 1904 [species]

Onthophagus planaticeps D'orbigny, 1913 [species]

Onthophagus planiceps Macleay, 1886 [species]

Onthophagus planicollis Harold, 1880 [species]

Onthophagus planifrons Frey, 1962 [species]

Onthophagus platalea Arrow, 1941 [species]

Onthophagus plato Bates, 1888 [species]

Onthophagus platypus Zhang, 1997 [species]

Onthophagus plebejus Klug, 1855 [species]

Onthophagus pleurogonus D'orbigny, 1913 [species]

Onthophagus plicatifrons D'orbigny, 1908 [species]

Onthophagus poeophagus Kabakov, 1979 [species]

Onthophagus poggii Scheuern, 1996 [species]

Onthophagus politissimus D'orbigny, 1908 [species]

Onthophagus politus Fabricius, 1798 [species]

Onthophagus pollicatus Harold, 1879 [species]

Onthophagus polyedrus D'orbigny, 1905 [species]

Onthophagus polyodon D'orbigny, 1913 [species]

Onthophagus polystigma D'orbigny, 1902 [species]

Onthophagus ponticus Harold, 1883 [species]

Onthophagus pooensis Cambefort & Nicolas, 1991 [species]

Onthophagus popovi Kabakov, 1983 [species]

Onthophagus porcus Arrow, 1931 [species]

Onthophagus poringensis Ochi & Kon, 2005 [species]

Onthophagus porrectus Reiche, 1847 [species]

Onthophagus possoi Walter, 1982 [species]

Onthophagus posticicornis D'orbigny, 1913 [species]

Onthophagus posticus Erichson, 1842 [species]

Onthophagus potanini Kabakov, 1979 [species]

O. p. hiurai

Onthophagus praecavatus D'Orbigny, 1913 [species]

Onthophagus praecellens Bates, 1887 [species]

Onthophagus praedatus Harold, 1862 [species]

Onthophagus praedentatus D'Orbigny, 1904 [species]

Onthophagus praefossus D'Orbigny, 1913 [species]

Onthophagus praelaminatus Frey, 1957 [species]

Onthophagus praestans Péringuey, 1901 [species]

Onthophagus praetortus Péringuey, 1904 [species]

Onthophagus prehensilis Arrow, 1920 [species]

Onthophagus privus Kabakov, 1998 [species]

Onthophagus probus Péringuey, 1901 [species]

Onthophagus procurvus Balthasar, 1935 [species]

Onthophagus producticollis d'Orbigny 1908 [species]

Onthophagus productus Arrow, 1907 [species]

Onthophagus profanus Frey, 1958 [species]

Onthophagus proletarius Harold, 1875 [species]

Onthophagus promontorii Boucomont, 1924 [species]

Onthophagus pronus Erichson, 1842 [species]

Onthophagus propinquus Macleay, 1888 [species]

Onthophagus propraecellens HOWDEN & GILL, 1987 [species]

Onthophagus prostans Reiche, 1847 [species]

Onthophagus proteus D'orbigny, 1902 [species]

Onthophagus protuberans Frey, 1955 [species]

Onthophagus proximus D'orbigny, 1908 [species]

Onthophagus pseudoaeneus D'orbigny, 1908 [species]

Onthophagus pseudoalcyon d'Orbigny 1913 [species]

Onthophagus pseudoarmatus Balthasar, 1944 [species]

Onthophagus pseudobidens D'orbigny, 1915 [species]

Onthophagus pseudobrutus Kabakov, 1983 [species]

Onthophagus pseudocaccobius Reitter, 1889 [species]

Onthophagus pseudocoracinus Kabakov, 1983 [species]

Onthophagus pseudofimetarius Balthasar, 1946 [species]

Onthophagus pseudoflexicollis Moretto, 2004 [species]

Onthophagus pseudohystrix Masumoto, 1995 [species]

Onthophagus pseudojaponicus Balthasar, 1941 [species]

O. p. fukiensis

Onthophagus pseudojavanus Paulian, 1931 [species]

Onthophagus pseudoliberianus Moretto, 2010 [species]

Onthophagus pseudopilosus Frey, 1958 [species]

Onthophagus pseudoplebejus Moretto, 2009 [species]

Onthophagus pseudorudis Ochi & Araya, 1992 [species]

Onthophagus pseudosanguineus WALTER & CAMBEFORT, 1977 [species]

Onthophagus pseudosellatus Balthasar, 1964 [species]

Onthophagus pseudoundulans ZUNINO & HALFFTER, 1988 [species]

O. p. howdeni

Onthophagus pseudovinctus Tagliaferri & Moretto, 2012 [species]

Onthophagus pseudovirens D'Orbigny, 1913 [species]

Onthophagus pseudoworoae Ochi, Kon & Barclay, 2009 [species]

Onthophagus psychopompus Ziani & Garakhloo, 2010 [species]

Onthophagus ptox Erichson, 1847 [species]

Onthophagus puberulus D'orbigny, 1902 [species]

Onthophagus pugionatus Fahraeus, 1857 [species]

Onthophagus pugnacior Blackburn, 1903 [species]

Onthophagus pugnax Harold, 1868 [species]

Onthophagus pulchellus D'orbigny, 1905 [species]

Onthophagus pullatus D'orbigny, 1905 [species]

Onthophagus pullus Roth, 1851 [species]

Onthophagus punctator Reitter, 1893 [species]

Onthophagus punctatus Illiger, 1803 [species]

Onthophagus punneeae Masumoto, 1989 [species]

Onthophagus punthari Storey, 1977 [species]

Onthophagus pupillatus Kolbe, 1886 [species]

Onthophagus purifrons D'orbigny, 1908 [species]

Onthophagus purpurascens Boucomont, 1914 [species]

Onthophagus purpureicollis Macleay, 1864 [species]

Onthophagus pusillus Fabricius, 1798 [species]

Onthophagus pusio Fahraeus, 1857 [species]

Onthophagus pygargus Motschulsky, 1845 [species]

Onthophagus pygidialis Lansberge, 1883 [species]

Onthophagus pygmaeus Schaller, 1783 [species]

Onthophagus pyramidalis Klug, 1855 [species]

Onthophagus quadraticeps Harold, 1867 [species]

Onthophagus quadriarmatus Fairmaire, 1892 [species]

Onthophagus quadricallosus D'orbigny, 1902 [species]

Onthophagus quadricolor Kabakov, 1983 [species]

Onthophagus quadricristatus D'Orbigny, 1908 [species]

Onthophagus quadricuspis d'Orbigny 1908 [species]

Onthophagus quadridentatus Fabricius, 1798 [species]

Onthophagus quadrilunatus D'orbigny, 1902 [species]

Onthophagus quadrimaculatus Raffray, 1877 [species]

Onthophagus quadrinodosus Fahraeus, 1857 [species]

Onthophagus quadrinodus Reitter, 1896 [species]

Onthophagus quadrinotatus D'orbigny, 1905 [species]

Onthophagus quadripustulatus Fabricius, 1775 [species]

Onthophagus quadrispinosus D'Orbigny, 1908 [species]

Onthophagus quadrituber D'Orbigny, 1908 [species]

Onthophagus quaestus Sharp, 1875 [species]

Onthophagus quasijohkii Ochi & Kon, 2005 [species]

Onthophagus quasitagal Ochi & Kon, 2005 [species]

Onthophagus queenslandicus Blackburn, 1903 [species]

Onthophagus quetzalis HOWDEN & GILL, 1993 [species]

Onthophagus quezonensis Ochi & Araya, 1992 [species]

Onthophagus quinquetuberculatus Macleay, 1871 [species]

Onthophagus quiproquo Moretto & Génier, 2010 [species]

Onthophagus rachelis Martin-Piera, 1985 [species]

Onthophagus ragazzii D'orbigny, 1904 [species]

Onthophagus rakovici Pittino, 2004 [species]

Onthophagus ramosellus Bates, 1891 [species]

Onthophagus ramosicornis d'Orbigny, 1902 [species]

Onthophagus ramosus Wiedemann, 1823 [species]

Onthophagus rana Arrow, 1931 [species]

Onthophagus rangifer Klug, 1855 [species]

Onthophagus ranongensis Masumoto, Ochi & Hanboonsong, 2008 [species]

Onthophagus ranunculus Arrow, 1913 [species]

Onthophagus rarus Guérin-Méneville, 1829 [species]

Onthophagus rasidorsis D'Orbigny, 1905 [species]

Onthophagus rasipennis D'orbigny, 1908 [species]

Onthophagus ratchasimaensis Masumoto & Hanboonsong & Ochi, 2002 [species]

Onthophagus rechingeri Petrovitz, 1980 [species]

Onthophagus rectecornutus Lansberge, 1883 [species]

Onthophagus rectefurcatus Fairmaire, 1891 [species]

Onthophagus rectestriatus D'orbigny, 1915 [species]

Onthophagus rectilamina D'orbigny, 1902 [species]

Onthophagus rectorispauliani Cambefort, 1980 [species]

Onthophagus refulgens Arrow, 1931 [species]

Onthophagus regalis Arrow, 1907 [species]

Onthophagus remotus Harold, 1879 [species]

Onthophagus renaudpauliani Ochi & Araya, 1996 [species]

Onthophagus reticollis Macleay, 1886 [species]

Onthophagus reticulatus D'orbigny, 1902 [species]

Onthophagus reticuliger Frey, 1960 [species]

Onthophagus revoili Lansberge, 1882 [species]

Onthophagus reyesi ZUNINO & HALFFTER, 1988 [species]

Onthophagus rhinocerus Gomes Alves, 1944 [species]

Onthophagus rhinolophus Harold, 1869 [species]

Onthophagus rhinophyllus Harold, 1868 [species]

Onthophagus rhodesianus Frey, 1975 [species]

Onthophagus rhombocephalus Kabakov, 1983 [species]

Onthophagus rhynchophorus Péringuey, 1904 [species]

Onthophagus ribbei Boucomont, 1914 [species]

Onthophagus riekoae Ochi & Kon, 2005 [species]

Onthophagus riparius Lansberge, 1885 [species]

Onthophagus ritsemai Lansberge, 1883 [species]

Onthophagus robertopoggii Ochi & Kon, 2006 [species]

Onthophagus rodentium Pittino, 2004 [species]

Onthophagus rorarius Harold, 1877 [species]

Onthophagus rosenbergi Krikken & Huijbregts, 2011 [species]

O. r. divergens

Onthophagus rosettae Frey, 1958 [species]

Onthophagus rostratus Harold, 1869 [species]

Onthophagus rotundatus D'orbigny, 1905 [species]

Onthophagus rotundibasis D'orbigny, 1902 [species]

Onthophagus rotundicollis Lansberge, 1883 [species]

Onthophagus roubali Balthasar, 1935 [species]

Onthophagus rougonorum Cambefort, 1984 [species]

Onthophagus rouyeri Boucomont, 1914 [species]

Onthophagus royi Biswas & Chatterjee, 1985 [species]

Onthophagus ruandanus Frey, 1971 [species]

Onthophagus rubefactus D'orbigny, 1913 [species]

Onthophagus rubellus D'orbigny, 1908 [species]

Onthophagus rubens D'orbigny, 1902 [species]

Onthophagus rubenticollis D'orbigny, 1902 [species]

Onthophagus rubescens Macleay, 1888 [species]

Onthophagus rubicundulus Macleay, 1871 [species]

Onthophagus rubidus D'orbigny, 1913 [species]

Onthophagus rubrescens Blanchard, 1843 [species]

Onthophagus rubricatus D'orbigny, 1902 [species]

Onthophagus rubricollis Hope, 1831 [species]

Onthophagus rubrimaculatus Macleay, 1864 [species]

Onthophagus rubripennis Arrow, 1907 [species]

Onthophagus rudis Sharp, 1875 [species]

Onthophagus rufescens Bates, 1887 [species]

Onthophagus ruficapillus Brullé, 1832 [species]

Onthophagus ruficauda Arrow, 1931 [species]

Onthophagus rufimanus Kabakov, 1982 [species]

Onthophagus rufiobscurior Ochi, Kon & Tsubaki, 2009 [species]

Onthophagus rufipodex D'Orbigny, 1913 [species]

Onthophagus rufobasalis Fairmaire, 1887 [species]

Onthophagus rufocastaneus Fairmaire, 1887 [species]

Onthophagus rufolimbatus D'orbigny, 1913 [species]

Onthophagus rufonotatus D'orbigny, 1902 [species]

Onthophagus rufopygus Frey, 1957 [species]

Onthophagus rufosignatus Macleay, 1864 [species]

Onthophagus rufostillans D'orbigny, 1907 [species]

Onthophagus rufovirens D'orbigny, 1904 [species]

Onthophagus rugicollis Harold, 1880 [species]

Onthophagus rugidorsis D'orbigny, 1913 [species]

Onthophagus rugipennis D'orbigny, 1908 [species]

Onthophagus rugosipennis Frey, 1973 [species]

Onthophagus rugosissimus D'orbigny, 1913 [species]

Onthophagus rugulipennis Fairmaire, 1887 [species]

Onthophagus rugulosus Harold, 1886 [species]

Onthophagus rupicapra Waterhouse, 1894 [species]

Onthophagus rutilans Sharp, 1875 [species]

O. r. aborneensis

Onthophagus sabahensis Ochi & Kon, 2006 [species]

Onthophagus sacharovskii Olsoufieff, 1918 [species]

Onthophagus saggitarius Fabricius, 1775 [species]

Onthophagus sagittarius Fabricius, 1775 [species]

Onthophagus sahai Biswas & Chatterjee, 1986 [species]

Onthophagus saïgonensis Boucomont, 1924 [species]

Onthophagus sakaeratensis Masumoto & Hanboonsong & Ochi, 2002 [species]

Onthophagus sakainoi Masumoto, 1991 [species]

Onthophagus sakamakii Ochi, Kon & Kawahara, 2010 [species]

Onthophagus salakensis Ochi, Kon & Hartini, 2010 [species]

Onthophagus salebrosus Macleay, 1888 [species]

Onthophagus saleyeri Lansberge, 1883 [species]

Onthophagus salvadorensis ZUNINO & HALFFTER, 1988 [species]

Onthophagus salvazai Paulian, 1978 [species]

Onthophagus samai Ziani, 2011 [species]

Onthophagus samoengus Masumoto, Ochi & Hanboonsong, 2008 [species]

Onthophagus sanggona Huijbregts & Krikken, 2009 [species]

Onthophagus sangirensis Boucomont, 1914 [species]

Onthophagus sanguineus D'orbigny, 1902 [species]

Onthophagus sanguinolentus D'orbigny, 1908 [species]

Onthophagus sangwalus Masumoto, Ochi & Hanboonsong, 2008 [species]

Onthophagus sansibaricus Harold, 1878 [species]

Onthophagus sapphirinus Fahraeus, 1857 [species]

Onthophagus sarasinorum Krikken & Huijbregts, 2011 [species]

Onthophagus sarawacus Harold, 1877 [species]

Onthophagus sasajii Ochi & Kon, 2001 [species]

Onthophagus satoi Ochi, Shimada & Kon, 2003 [species]

Onthophagus saudiensis Frey, 1962 [species]

Onthophagus sauteri Gillet, 1924 [species]

Onthophagus savanicola Cambefort, 1984 [species]

Onthophagus sayapensis Ochi & Kon, 2005 [species]

Onthophagus scaber Roth, 1851 [species]

Onthophagus scaberrimus D'orbigny, 1904 [species]

Onthophagus scabriusculus Harold, 1873 [species]

Onthophagus scapularis D'orbigny, 1902 [species]

Onthophagus sceptrifer Boucomont, 1924 [species]

Onthophagus schaefernai Balthasar, 1935 [species]

Onthophagus schaufussi Harold, 1867 [species]

Onthophagus schawalleri Scheuern, 1996 [species]

Onthophagus schillhammeri Kabakov, 2006 [species]

Onthophagus schoolmeestersi Ochi & Kon, 2007 [species]

Onthophagus schunckei Paulian, 1936 [species]

Onthophagus schwaneri Snellen van Vollenhoven, 1864 [species]

Onthophagus sciron Balthasar, 1969 [species]

Onthophagus scotti Masumoto, Ochi & Hanboonsong, 2002 [species]

Onthophagus scrutator Harold, 1877 [species]

Onthophagus sculptilis Gerstaecker, 1871 [species]

Onthophagus seabrai Gomes Alves, 1944 [species]

Onthophagus secundarius Roth, 1851 [species]

Onthophagus sellatulus D'orbigny, 1902 [species]

Onthophagus sellatus Klug, 1845 [species]

Onthophagus sembeli Krikken & Huijbregts, 2008 [species]

Onthophagus semiaratus D'orbigny, 1902 [species]

Onthophagus semiasper D'orbigny, 1902 [species]

Onthophagus semiaureus Lansberge, 1883 [species]

Onthophagus semichalcites D'orbigny, 1902 [species]

Onthophagus semicínctus D'Orbigny, 1897 [species]

Onthophagus semicornis Panzer, 1798 [species]

Onthophagus semicroceus D'orbigny, 1915 [species]

Onthophagus semicupreus Harold, 1877 [species]

Onthophagus semidanumensis Ochi, Kon & Barclay, 2009 [species]

Onthophagus semifex Krikken & Huijbregts, 2008 [species]

Onthophagus semiflavus Boheman, 1860 [species]

Onthophagus semigraniger D'orbigny, 1905 [species]

Onthophagus semigranosus Lansberge, 1883 [species]

Onthophagus semiiris Thomson, 1858 [species]

Onthophagus semilaevis D'orbigny, 1913 [species]

Onthophagus semimetallicus Lea, 1923 [species]

Onthophagus seminitens D'orbigny, 1902 [species]

Onthophagus seminitidus D'orbigny, 1908 [species]

Onthophagus semiopacus Harold, 1869 [species]

Onthophagus semipacificus Ochi & Kon, 2006 [species]

Onthophagus semipersonatus Ochi, Kon & Tsubaki, 2009 [species]

Onthophagus semipiceus Kabakov, 1998 [species]

Onthophagus semivestitus Paulian, 1937 [species]

Onthophagus semivirescens D'orbigny, 1902 [species]

Onthophagus semiviridis D'orbigny, 1904 [species]

Onthophagus senegalensis D'orbigny, 1902 [species]

Onthophagus senescens Péringuey, 1908 [species]

Onthophagus senex Boucomont, 1914 [species]

Onthophagus seniculus Fabricius, 1781 [species]

Onthophagus sepilokensis Ochi & Kon, 2006 [species]

Onthophagus seramicus Huijbregts & Krikken, 2012 [species]

Onthophagus serapiensis Ochi, Kon & Barclay, 2008 [species]

Onthophagus serdangensis Lansberge, 1886 [species]

Onthophagus sericans Frey, 1975 [species]

Onthophagus sericatus Reitter, 1893 [species]

Onthophagus sericeicollis Boucomont, 1928 [species]

Onthophagus serienotatus D'orbigny, 1902 [species]

Onthophagus setchan Masumoto, 1984 [species]

Onthophagus setoculus Krikken & Huijbregts, 1987 [species]

Onthophagus setosus Fahraeus, 1857 [species]

Onthophagus sexcornutus d'Orbigny, 1902 [species]

Onthophagus sexdentatus Boucomont, 1919 [species]

Onthophagus sexstriatus Montrouzier, 1855 [species]

Onthophagus shanicus Kabakov, 2006 [species]

Onthophagus shapovalovi Gusakov, 2012 [species]

Onthophagus sharpi Harold, 1875 [species]

Onthophagus shibatai Nakane, 1960 [species]

Onthophagus shillongensis Scheuern, 1995 [species]

Onthophagus shimba Cambefort, 1980 [species]

Onthophagus shirakii Nakane, 1960 [species]

Onthophagus shizumui Ochi & Araya, 1992 [species]

Onthophagus sibela Huijbregts & Krikken, 2012 [species]

Onthophagus sibiricus Harold, 1877 [species]

Onthophagus sibuyanus Boucomont, 1924 [species]

Onthophagus sidama Gestro, 1895 [species]

Onthophagus sideki Krikken & Huijbregts, 1987 [species]

Onthophagus signaticollis Frey, 1970 [species]

Onthophagus signatus Fahraeus, 1857 [species]

Onthophagus signifer Harold, 1877 [species]

Onthophagus sihkahonoi Huijbregts & Krikken, 2009 [species]

Onthophagus sikkimensis Gillet, 1925 [species]

Onthophagus silus Balthasar, 1960 [species]

Onthophagus simboroni Ochi & Kon, 2006 [species]

Onthophagus similis Scriba 1790 [species]

Onthophagus simillimus D'orbigny, 1913 [species]

Onthophagus simius Reitter, 1893 [species]

Onthophagus simoni D'orbigny, 1902 [species]

Onthophagus simplex Raffray, 1877 [species]

Onthophagus simpliciceps D'orbigny, 1905 [species]

Onthophagus simplicifrons Reitter, 1893 [species]

Onthophagus simulator D'orbigny, 1905 [species]

Onthophagus sinagai Krikken & Huijbregts, 2008 [species]

Onthophagus singhaakhomus Masumoto, 1992 [species]

Onthophagus singulariformis Kohlmann & Solis, 2001 [species]

Onthophagus sinicus Zhang & Wang, 1997 [species]

Onthophagus sinuosicollis D'orbigny, 1905 [species]

Onthophagus sinuosus D'orbigny, 1913 [species]

Onthophagus sipilouensis Cambefort, 1984 [species]

Onthophagus sisyphoides Krikken, 1977 [species]

Onthophagus sjoestethi D'Orbigny, 1904 [species]

Onthophagus sloanei Blackburn, 1903 [species]

Onthophagus smeenki Cambefort, 1980 [species]

Onthophagus smetanai Balthasar, 1952 [species]

Onthophagus snoflaki Balthasar, 1944 [species]

Onthophagus sobrius Balthasar, 1959 [species]

Onthophagus socialis Arrow 1931 [species]

Onthophagus solidus Gillet, 1927 [species]

Onthophagus solisi HOWDEN & GILL, 1993 [species]

Onthophagus solivagus Harold, 1886 [species]

Onthophagus solmani Stebnicka, 1975 [species]

Onthophagus somalicola Balthasar, 1941 [species]

Onthophagus somalicus d'Orbigny, 1902 [species]

Onthophagus sonani Miwa, 1930 [species]

Onthophagus songsokensis Biswas & Chatterjee, 1985 [species]

Onthophagus sophiae Nicolas, 2006 [species]

Onthophagus sparsepunctatus Frey, 1956 [species]

Onthophagus sparsulus Reitter, 1893 [species]

Onthophagus spathatus Boucomont, 1914 [species]

Onthophagus speculatus D'Orbigny, 1908 [species]

Onthophagus speculicollis Quedenfeldt 1884 [species]

Onthophagus speculifer Solsky, 1876 [species]

Onthophagus spiculatus Boucomont, 1914 [species]

Onthophagus spinicornis Gillet, 1930 [species]

Onthophagus spinifex Fabricius, 1781 [species]

Onthophagus splendidus Boucomont, 1932 [species]

Onthophagus spurcatus D'orbigny, 1905 [species]

Onthophagus squalidus Lea, 1923 [species]

Onthophagus stanleyi Moretto & Génier, 2010 [species]

Onthophagus stehliki Balthasar, 1966 [species]

Onthophagus steinheili Harold, 1880 [species]

Onthophagus stellio Erichson, 1843 [species]

Onthophagus stenocerus Harold, 1867 [species]

Onthophagus sternalis Arrow, 1931 [species]

Onthophagus sternax Balthasar, 1959 [species]

Onthophagus sticticus Harold, 1867 [species]

Onthophagus stictus D'orbigny, 1913 [species]

Onthophagus stigmosus D'orbigny, 1902 [species]

Onthophagus stillatus D'orbigny, 1904 [species]

Onthophagus stockwelli Howden & Young, 1981 [species]

Onthophagus stomachosus Krell, 2009 [species]

Onthophagus strabo Reitter 1892 [species]

Onthophagus strandi Balthasar, 1935 [species]

Onthophagus streltsovi Tarasov & Kabakov, 2010 [species]

Onthophagus striaticollis Kabakov, 1983 [species]

Onthophagus strictestriatus D'orbigny, 1913 [species]

Onthophagus strnadi Kabakov, 1998 [species]

Onthophagus stuhlmanni D'orbigny, 1908 [species]

Onthophagus stylocerus Graells, 1851 [species]

Onthophagus subalternans D'orbigny, 1902 [species]

Onthophagus subansiriensis Biswas, 1979 [species]

Onthophagus subcancer Howden, 1973 [species]

Onthophagus subcinctus D'orbigny, 1913 [species]

Onthophagus subcornutus Boucomont, 1914 [species]

Onthophagus subdivisus D'orbigny, 1908 [species]

Onthophagus subextensus Kolbe, 1895 [species]

Onthophagus subglaber D'orbigny, 1902 [species]

Onthophagus subhumeralis D'orbigny, 1902 [species]

Onthophagus subnudus D'orbigny, 1902 [species]

Onthophagus subocellatus D'orbigny, 1905 [species]

Onthophagus subplanus D'orbigny, 1902 [species]

Onthophagus subrugosus D'orbigny, 1905 [species]

Onthophagus subsapaensis Kabakov, 1994 [species]

Onthophagus subsulcatus D'orbigny, 1908 [species]

Onthophagus subulifer D'orbigny, 1908 [species]

Onthophagus suermelii Petrovitz 1963 [species]

Onthophagus suffusus Klug, 1855 [species]

Onthophagus sugihartoi Ochi, 2007 [species]

Onthophagus sugillatus Klug, 1855 [species]

Onthophagus suginoi Ochi, 1984 [species]

Onthophagus suginokoichii Ochi & Kon, 2008 [species]

Onthophagus suillus Arrow, 1931 [species]

Onthophagus sulawesiensis Krikken & Huijbregts, 2011 [species]

Onthophagus sulcatulus D'orbigny, 1907 [species]

Onthophagus sulci Balthasar, 1935 [species]

Onthophagus sulcipennis D'orbigny, 1902 [species]

Onthophagus sumatramontanus Ochi & Kon, 2008 [species]

Onthophagus sumatranus Lansberge, 1883 [species]

Onthophagus sumawacus Krikken & Huijbregts, 2008 [species]

Onthophagus sumbavensis Boucomont, 1914 [species]

Onthophagus sumptuosus Paulian, 1937 [species]

Onthophagus sunantaae Masumoto, 1989 [species]

Onthophagus sundanensis Lansberge, 1883 [species]

Onthophagus superbus D'Orbigny, 1904 [species]

Onthophagus surdus Boucomont, 1925 [species]

Onthophagus susterai Balthasar, 1952 [species]

Onthophagus sutiliceps D'Orbigny, 1902 [species]

Onthophagus sutleinensis Splichal, 1910 [species]

Onthophagus suturalis Péringuey, 1888 [species]

Onthophagus suturellus Brullé, 1832 [species]

Onthophagus sycophanta Fairmaire, 1887 [species]

Onthophagus sydneyensis Blackburn, 1903 [species]

Onthophagus sylvestris WALTER & CAMBEFORT, 1977 [species]

Onthophagus symbioticus Arrow, 1920 [species]

Onthophagus synceri Cambefort, 1984 [species]

Onthophagus taayai Masumoto, 1995 [species]

Onthophagus tabellicornis Macleay, 1864 [species]

Onthophagus tabellifer Gillet, 1927 [species]

Onthophagus tabidus Balthasar, 1935 [species]

Onthophagus taboranus D'orbigny, 1902 [species]

Onthophagus taeniatus Boucomont, 1914 [species]

Onthophagus tagal Boucomont, 1924 [species]

Onthophagus taichii Ochi, Kon & Barclay, 2008 [species]

Onthophagus taiensis Cambefort, 1984 [species]

Onthophagus taiwanus Nomura, 1973 [species]

Onthophagus taiyaruensis Masumoto, 1977 [species]

Onthophagus takakui Ochi, Kon & Hartini, 2009 [species]

Onthophagus takedai Ochi & Araya, 1992 [species]

Onthophagus takeshii Ochi & Araya, 1992 [species]

Onthophagus talpa Fahraeus, 1857 [species]

Onthophagus tambing Krikken & Huijbregts, 2011 [species]

Onthophagus tamijii Kon, Sakai & Ochi, 2000 [species]

Onthophagus tamworthi Blackburn, 1903 [species]

Onthophagus tanganus D'orbigny, 1913 [species]

Onthophagus taoi Ochi & Kon, 2006 [species]

Onthophagus tapirus Sharp, 1877 [species]

Onthophagus taprobanus Arrow, 1931 [species]

Onthophagus tarandus Fabricius, 1792 [species]

Onthophagus tarascus ZUNINO & HALFFTER, 1988 [species]

O. t. jaliscensis

Onthophagus tarsius Arrow, 1941 [species]

Onthophagus taruni Biswas & Chatterjee, 1986 [species]

Onthophagus tatsienluensis Balthasar, 1942 [species]

Onthophagus taurinus White, 1844 [species]

Onthophagus tauroides Gillet, 1930 [species]

Onthophagus taxillus Balthasar, 1969 [species]

Onthophagus taymansi D'Orbigny, 1908 [species]

Onthophagus teitanicus D'orbigny, 1902 [species]

Onthophagus telegonus Balthasar, 1969 [species]

Onthophagus telephus Balthasar, 1969 [species]

Onthophagus temporalis D'orbigny, 1902 [species]

Onthophagus tenax Balthasar, 1964 [species]

Onthophagus tenebrosus Harold, 1871 [species]

Onthophagus tenuigraniger D'orbigny, 1913 [species]

Onthophagus tenuistriatus D'orbigny, 1905 [species]

Onthophagus terminatus Eschscholtz, 1822 [species]

Onthophagus terrara Storey, 1977 [species]

Onthophagus tersicollis Müller, 1947 [species]

Onthophagus tersidorsis D'Orbigny, 1902 [species]

Onthophagus tersipennis D'orbigny, 1902 [species]

Onthophagus tersus D'orbigny, 1913 [species]

Onthophagus tesquorum Semenov & Medvedev 1927 [species]

Onthophagus tesseratus D'orbigny, 1908 [species]

Onthophagus tessulatus Harold, 1871 [species]

Onthophagus testaceoviolaceus Paulian, 1937 [species]

Onthophagus tetricus Harold, 1877 [species]

Onthophagus thai Kabakov, 1994 [species]

Onthophagus thailaevis Masumoto, Ochi & Hanboonsong, 2008 [species]

Onthophagus thainuaensis Masumoto, Ochi & Hanboonsong, 2008 [species]

Onthophagus thaitai Masumoto, Ochi & Hanboonsong, 2008 [species]

Onthophagus thanwaakhomus Masumoto, 1992 [species]

Onthophagus tholaayi Masumoto, 1990 [species]

Onthophagus thoreyi Harold, 1868 [species]

Onthophagus tiamicus Kabakov, 1994 [species]

Onthophagus tibetanus Arrow, 1907 [species]

Onthophagus tibialis Frey, 1963 [species]

O. t. diversiceps

Onthophagus tigrinus D'orbigny, 1908 [species]

Onthophagus timorensis Boucomont, 1914 [species]

Onthophagus tiniocelloides Boucomont, 1925 [species]

Onthophagus tirapensis Biswas & Chatterjee, 1985 [species]

Onthophagus togeman Matthews, 1972 [species]

Onthophagus tongbantumi Masumoto, Ochi & Hanboonsong, 2002 [species]

Onthophagus tonkineus Gillet, 1921 [species]

Onthophagus tonsus D'orbigny, 1902 [species]

Onthophagus tonywhitteni Krikken & Huijbregts, 2011 [species]

Onthophagus toraut Krikken & Huijbregts, 2011 [species]

Onthophagus totonicapamus Bates, 1887 [species]

Onthophagus traginus Frey, 1972 [species]

Onthophagus tragoides Boucomont, 1914 [species]

Onthophagus tragus Fabricius, 1792 [species]

Onthophagus transisthmius Howden & Young, 1981 [species]

Onthophagus transquadridentatus Scheuern, 1995 [species]

Onthophagus transvestitus Huijbregts & Krikken, 2009 [species]

Onthophagus trapezicornis D'orbigny, 1902 [species]

Onthophagus traversii D'orbigny, 1904 [species]

Onthophagus trawalla STOREY & WEIR, 1990 [species]

Onthophagus triacanthus Castelnau, 1840 [species]

Onthophagus trianguliceps D'Orbigny, 1905 [species]

Onthophagus triarmatus D'Orbigny, 1904 [species]

Onthophagus tricariniger D'orbigny, 1902 [species]

Onthophagus tricavicollis Lea, 1923 [species]

Onthophagus triceratops Arrow 1913 [species]

Onthophagus trichopygus D'orbigny, 1905 [species]

Onthophagus tricolor Boucomont, 1914 [species]

Onthophagus tricornifrons D'Orbigny, 1908 [species]

Onthophagus tricorniger Boheman, 1860 [species]

Onthophagus tricornis Wiedemann, 1823 [species]

Onthophagus tridenticeps D'orbigny, 1902 [species]

Onthophagus tridentitibialis Ochi & Kon, 2008 [species]

Onthophagus trifidisetis D'orbigny, 1905 [species]

Onthophagus trigibber Reitter, 1893 [species]

Onthophagus triimpressus D'orbigny, 1905 [species]

Onthophagus trinodosus Fahraeus, 1857 [species]

Onthophagus trinominatus Goidanich, 1926 [species]

Onthophagus tripartitus D'orbigny, 1902 [species]

Onthophagus tripolitanus Heyden, 1890 [species]

Onthophagus triptolemus Balthasar, 1969 [species]

Onthophagus trispinus Reitter, 1892 [species]

Onthophagus tristis Harold, 1873 [species]

Onthophagus tritinctus Boucomont, 1914 [species]

Onthophagus trituber Wiedemann, 1823 [species]

O. t. jacobsoni

Onthophagus triundulatus Balthasar, 1964 [species]

Onthophagus trochilus Arrow, 1931 [species]

Onthophagus troglodyta Wiedemann, 1823 [species]

Onthophagus troniceki Balthasar, 1933 [species]

Onthophagus truchmenus Kolenati, 1846 [species]

O. t. iranicus, O. t. turkmeniacus

Onthophagus truncaticornis Schaller, 1783 [species]

Onthophagus trungboensis Kabakov, 1983 [species]

Onthophagus tschadensis Balthasar, 1963 [species]

Onthophagus tschoffeni D'orbigny, 1904 [species]

Onthophagus tshuapae Balthasar, 1964 [species]

Onthophagus tsubakii Ochi & Kon, 2009 [species]

Onthophagus tubericollis Raffray, 1877 [species]

Onthophagus tuckonie Matthews, 1972 [species]

Onthophagus tumami Masumoto & Ochi & Hanboonsong, 2002 [species]

Onthophagus tumidulus Gerstaecker, 1871 [species]

Onthophagus tungkamangensis Masumoto, Ochi & Hanboonsong, 2007 [species]

Onthophagus tungkamungensis Masumoto, Ochi & Hanboonsong, 2008 [species]

Onthophagus turbatus Walker, 1858 [species]

Onthophagus turfanicus Balthasar, 1971 [species]

Onthophagus turgidus Kohlmann & Solis, 2012 [species]

Onthophagus turpidoides Kabakov, 2008 [species]

Onthophagus turpidus Reitter, 1887 [species]

Onthophagus turrbal Matthews, 1972 [species]

Onthophagus tuzetae WALTER & CAMBEFORT, 1977 [species]

Onthophagus tweedensis Blackburn, 1903 [species]

Onthophagus uedai Ochi & Kon, 2006 [species]

Onthophagus uelensis Frey, 1960 [species]

Onthophagus uenoi Ochi 1995 [species]

O. u. borneomontanus

Onthophagus ugandicus Frey, 1975 [species]

Onthophagus ukerewensis Balthasar, 1937 [species]

Onthophagus uluguru Frey, 1975 [species]

Onthophagus ulula Balthasar, 1966 [species]

Onthophagus umbilicatus D'orbigny, 1908 [species]

Onthophagus umbilicopunctatus Medvedev, 1959 [species]

Onthophagus umbratus D'orbigny, 1902 [species]

Onthophagus undaticeps D'orbigny, 1902 [species]

Onthophagus undulans Bates, 1887 [species]

O. u. oaxacanus

Onthophagus unguiculatus Kabakov, 1994 [species]

Onthophagus unicarina D'orbigny, 1904 [species]

Onthophagus unidens Quedenfeldt, 1884 [species]

Onthophagus unidentatus Boucomont, 1927 [species]

Onthophagus unifasciatus Schaller, 1783 [species]

Onthophagus uniformis Heyden, 1886 [species]

Onthophagus urellus Boucomont, 1919 [species]

Onthophagus ursinus D'orbigny, 1902 [species]

Onthophagus ursus Boucomont, 1926 [species]

Onthophagus usambaricus Paulian, 1937 [species]

Onthophagus usurpator Balthasar, 1959 [species]

Onthophagus utsunomiyae Masumoto, Ochi & Hanboonsong, 2007 [species]

Onthophagus vacca Linnaeus, 1767 [species]

Onthophagus vanasseni Krikken & Huijbregts, 2008 [species]

Onthophagus vanderblomi Krikken & Huijbregts, 2008 [species]

Onthophagus vaneyeni Frey, 1960 [species]

Onthophagus vanofwegeni Krikken & Huijbregts, 2008 [species]

Onthophagus varianus Lea, 1923 [species]

Onthophagus variatus D'orbigny, 1902 [species]

Onthophagus variegatus Fabricius, 1798 [species]

Onthophagus variegranosus D'orbigny, 1905 [species]

Onthophagus variolaris Lansberge, 1883 [species]

Onthophagus variolicollis Lea, 1923 [species]

Onthophagus variolosus D'orbigny, 1902 [species]

Onthophagus varius D'orbigny, 1913 [species]

Onthophagus vasiljevi Kabakov, 1983 [species]

Onthophagus vassei D'orbigny, 1908 [species]

Onthophagus vatovai Müller, 1942 [species]

Onthophagus vaulogeri Boucomont, 1923 [species]

Onthophagus velliger D'orbigny, 1907 [species]

Onthophagus ventralis Lansberge, 1883 [species]

Onthophagus ventrosus D'orbigny, 1905 [species]

Onthophagus venzoi Balthasar, 1941 [species]

Onthophagus veracruzensis Delgado & Pensado, 1998 [species]

Onthophagus vermiculatus Frey, 1970 [species]

Onthophagus verrucosus D'orbigny, 1902 [species]

Onthophagus versus D'Orbigny, 1913 [species]

Onthophagus versutus Péringuey, 1901 [species]

Onthophagus verticalis Fahraeus, 1857 [species]

Onthophagus verticicornis Laicharting, 1781 [species]

Onthophagus vesanus Balthasar, 1967 [species]

Onthophagus vespertilio Howden & Cartwright & Hallfter, 1956 [species]

Onthophagus vestitus D'orbigny, 1902 [species]

Onthophagus vethi Krikken, 1977 [species]

Onthophagus vicinus Harold, 1886 [species]

Onthophagus victor Sharp, 1875 [species]

Onthophagus victoriensis Blackburn, 1903 [species]

Onthophagus viduus Harold, 1875 [species]

Onthophagus vigens Péringuey, 1901 [species]

Onthophagus vigilans Boucomont, 1921 [species]

Onthophagus vilis Harold, 1877 [species]

Onthophagus villanuevai Delgado & Deloya, 1990 [species]

Onthophagus villosus Macleay, 1888 [species]

Onthophagus vinctoides Frey, 1957 [species]

Onthophagus vinctus Erichson, 1843 [species]

Onthophagus violaceotinctus Gillet, 1925 [species]

Onthophagus violaceoviridis Paulian, 1937 [species]

Onthophagus violetae ZUNINO & HALFFTER, 1997 [species]

Onthophagus virens d'Orbigny 1913 [species]

Onthophagus virescens Harold, 1867 [species]

Onthophagus viridiaereus D'orbigny, 1908 [species]

Onthophagus viridicatus D'orbigny, 1902 [species]

Onthophagus viridiceps d'Orbigny, 1910 [species]

Onthophagus viridicervicapra Ochi, Kon & Tsubaki, 2009 [species]

Onthophagus viridis Ménétriès, 1832 [species]

Onthophagus viriditinctus Reitter, 1893 [species]

Onthophagus viridivinosus Kohlmann & Solis, 2001 [species]

Onthophagus vitalisi Boucomont, 1921 [species]

Onthophagus vitulus Fabricius, 1776 [species]

Onthophagus vividus Arrow, 1907 [species]

Onthophagus viviensis D'orbigny, 1905 [species]

Onthophagus vladimiri Frey, 1957 [species]

Onthophagus vlasovi Medvedev, 1958 [species]

Onthophagus volucer Balthasar, 1959 [species]

Onthophagus vuattouxi Cambefort, 1984 [species]

Onthophagus vulpes Harold, 1877 [species]

Onthophagus vulpinaris Schönfeldt, 1906 [species]

Onthophagus vulpinus Arrow, 1931 [species]

Onthophagus vultuosus D'orbigny, 1902 [species]

Onthophagus vultur Arrow, 1931 [species]

Onthophagus vylderi D'orbigny, 1913 [species]

Onthophagus wagamen Matthews, 1972 [species]

Onthophagus wakelbura Matthews, 1972 [species]

Onthophagus wallacei Harold, 1871 [species]

Onthophagus walteri Macleay, 1887 [species]

Onthophagus waminda Matthews, 1972 [species]

Onthophagus wanappe Storey, 1977 [species]

Onthophagus wangi Masumoto, Chen & Ochi, 2004 [species]

Onthophagus wangnamkhieoensis Masumoto & Hanboonsong & Ochi, 2002 [species]

Onthophagus wangnamkieous Masumoto, Ochi & Hanboonsong, 2008 [species]

Onthophagus watanabei Ochi & Kon, 2002 [species]

Onthophagus waterhousei Gillet, 1927 [species]

Onthophagus waterloti D'orbigny, 1908 [species]

Onthophagus waterstradti Boucomont, 1914 [species]

Onthophagus watuwila Krikken & Huijbregts, 2008 [species]

Onthophagus wayaua Huijbregts & Krikken, 2012 [species]

Onthophagus wensis Josso & Prévost, 2006 [species]

Onthophagus weringerong STOREY & WEIR, 1990 [species]

Onthophagus werneri Moretto & Nicolas, 2002 [species]

Onthophagus wiebesi Krikken, 1977 [species]

Onthophagus wigmungan Matthews, 1972 [species]

Onthophagus wilgi Matthews, 1972 [species]

Onthophagus willameorum WALTER & CAMBEFORT, 1977 [species]

Onthophagus williamsi STOREY & WEIR, 1990 [species]

Onthophagus witteianus Krell, 2009 [species]

Onthophagus wombalano Matthews, 1972 [species]

Onthophagus woroae Ochi & Kon, 2006 [species]

O. w. bruneiensis

Onthophagus worooa STOREY & WEIR, 1990 [species]

Onthophagus worsissa Roth, 1851 [species]

Onthophagus xanthochlorus WALTER & CAMBEFORT, 1977 [species]

Onthophagus xanthomerus Bates, 1887 [species]

Onthophagus xanthopterus D'orbigny, 1908 [species]

Onthophagus xanthopygus D'orbigny, 1908 [species]

Onthophagus xiphias Solis & Kohlmann, 2003 [species]

Onthophagus yackatoon STOREY & WEIR, 1990 [species]

Onthophagus yakuinsulanus Nakane, 1984 [species]

Onthophagus yamaokai Masumoto & Ochi & Hanboonsong, 2002 [species]

Onthophagus yangi Masumoto, Tsai & Ochi, 2006 [species]

Onthophagus yanoi Matsumura, 1938 [species]

Onthophagus yaran STOREY & WEIR, 1990 [species]

Onthophagus yarrumba Storey, 1977 [species]

Onthophagus yeyeko Matthews, 1972 [species]

Onthophagus yiryoront Matthews, 1972 [species]

Onthophagus yourula STOREY & WEIR, 1990 [species]

Onthophagus yubarinus Matsumura, 1937 [species]

Onthophagus yucatanus Delgado-Castillo, Peraza & Deloya, 2006 [species]

Onthophagus yukae Masumoto, Ochi & Hanboonsong, 2002 [species]

Onthophagus yumotoi Ochi & Kon, 2006 [species]

Onthophagus yungaburra Matthews, 1972 [species]

Onthophagus yunkara Matthews, 1972 [species]

Onthophagus yunnanus Boucomont, 1912 [species]

Onthophagus yvescambeforti Moretto, 2004 [species]

Onthophagus zagrosicus Kabakov, 2006 [species]

Onthophagus zairensis WALTER & CAMBEFORT, 1977 [species]

Onthophagus zapotecus ZUNINO & HALFFTER, 1988 [species]

Onthophagus zavattarii Müller, 1939 [species]

Onthophagus zavreli Balthasar, 1935 [species]

Onthophagus zebra Arrow, 1931 [species]

Onthophagus zebu Boucomont, 1921 [species]

Onthophagus zetteli Masumoto & Ochi & Hanboonsong, 2002 [species]

Onthophagus zicsii Balthasar, 1967 [species]

Onthophagus zimmermanni Balthasar, 1959 [species]

Onthophagus zumpti Frey, 1954 [species]

Onthophagus zuninoi Martin-Piera, 1985 [species]

Onthophagus zymoticus Moxey, 1963 [species]

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