Copris megaceratoides Waterhouse, 1891

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Copris acutidens Motschulsky, 1860

Copris afgoi Cambefort, 1992

Copris agnus Sharp, 1875

Copris algol NGUYEN-PHUNG, 1989

Copris amabilis Kolbe, 1914

Copris amyntor Klug, 1855

Copris anceus Olivier, 1789

Copris andrewesi Waterhouse, 1891

Copris angusticornis Arrow, 1933

Copris angustus Nguyen-phung & Cambefort, 1986

Copris anomiopseoides Boucomont, 1924

Copris antares Ferreira, 1958

Copris apicepunctatus Balthasar, 1942

Copris arcturus Gillet, 1907

Copris ares Zelenka, 1993

Copris arimotoi OCHI & ARAYA, 1992

Copris arizonensis Schaeffer, 1906

Copris armatus Harold, 1869

Copris armeniacus Faldermann, 1835

Copris armiger Gillet, 1910

Copris arrowi Felsche, 1910

Copris aspericollis Gillet, 1910

Copris atropolitus Gillet, 1910

Copris barclayi Ochi, Kon & Bai, 2009

Copris basilewskyi Ferreira, 1962

Copris basipunctatus Balthasar, 1942

Copris bellator Chevrolat, 1844

Copris bengalensis Gillet, 1911

Copris bidens Kolbe, 1893

Copris bihamatus Balthasar, 1965

Copris bootes Klug, 1855

Copris boucardi Harold, 1869

Copris bovinus Gillet, 1908

Copris brachypterus Nomura, 1964

Copris brittoni Ferreira, 1962

Copris caelatus Fabricius, 1794

Copris caliginosus Kohlmann, Cano & Delgado, 2003

Copris cambeforti NGUYEN-PHUNG, 1988

Copris cambodiensis Ochi, Kon & Kawahara, 2008

Copris camerunus Felsche, 1904

Copris capensis Waterhouse, 1891

Copris cariniceps Felsche, 1910

Copris carinicus Gillet, 1910

Copris carmelita Fabricius, 1801

Copris cassius Péringuey, 1901

Copris celebensis Lansberge, 1886

Copris cheni Ochi, Kon & Bai, 2007

Copris colmanti Gillet, 1908

Copris complexus NGUYEN-PHUNG, 1989

Copris compressipennis Gillet, 1910

Copris confucius Harold, 1877

Copris coriarius Gillet, 1907

Copris cornifrons Boheman, 1860

Copris corniger Sahlberg, 1823

Copris corpulentus Gillet, 1910

Copris costaricensis Gahan, 1894

Copris cribratus Gillet, 1927

Copris cribricollis Gillet, 1910

Copris curlettii Moretto, 2011

Copris davisi Nguyen-phung & Cambefort, 1986

Copris davisoni Waterhouse, 1891

Copris decellei Cambefort, 1992

Copris delicatus Arrow, 1931

Copris denticulatus NGUYEN-PHUNG, 1988

Copris diversus Waterhouse, 1891

Copris doriae Harold, 1877

Copris draco Arrow, 1906

Copris dracunculus Fereira, 1959

Copris dudleyi Cambefort, 1992

Copris eburneus Cambefort, 1992

Copris elpheneroides Felsche, 1910

Copris elphenor Klug, 1855

Copris erratus Lansberge, 1886

Copris evanidus Klug, 1855

Copris excisus Waterhouse, 1891

Copris fallaciosus Gillet, 1907

Copris fallax Felsche, 1910

Copris felschei Reitter, 1893

Copris fidius Olivier, 1789

Copris frankenbergerianus Balthasar, 1958

Copris freyi Ferreira, 1961

Copris fricator Fabricius, 1787

Copris fujiokai Ochi & Kon, 1996

Copris fukiensis Balthasar, 1952

Copris furciceps Felsche, 1910

Copris garambae Cambefort, 1992

Copris gazellarum Gillet, 1918

Copris gibbulus Lansberge, 1886

Copris gilleti Kolbe, 1907

Copris gladiator Arrow, 1933

Copris gopheri Hubbard, 1894

Copris gracilis Waterhouse, 1891

Copris graueri Kolbe, 1914

Copris halffteri Matthews, 1959

Copris harrisi Waterhouse, 1891

Copris hartli Frey, 1975

Copris hatayamai OCHI & ARAYA, 1992

Copris hidakai Ochi & Kon, 1996

Copris hispanus (Linnaeus, 1764)

Copris howdeni MATTHEWS & HALFFTER, 1959

Copris humilis Gillet, 1908

Copris hybridus Gillet, 1910

Copris igualensis Warner, 1990

Copris illotus Balthasar, 1942

Copris imitans Felsche, 1910

Copris inaequabilis Zhang, 1997

Copris incertus Say, 1835

Copris inemarginatus Blatchley, 1918

Copris inhalatus Quedenfeldt, 1884

Copris insidiosus Péringuey, 1901

Copris integer Reiche, 1847

Copris interioris Kolbe, 1897

Copris iris Sharp, 1875

Copris jacchoides Nguyen-phung & Cambefort, 1987

Copris jacchus Fabricius, 1775

Copris jahi Nguyen-phung & Cambefort, 1986

Copris jucundus Gillet, 1932

Copris kachinensis Ochi, Kon & Kawahara, 2008

Copris kasagii Ochi & Kon, 1996

Copris katangae Cambefort & Nguyen-phung, 1996

Copris keralensis Gill, 1986

Copris kiuchii Masumoto, 1989

Copris klapperichi Balthasar, 1942

Copris klugi Harold, 1869

Copris kusakabei Ochi, Kon & Kawahara, 2005

Copris laeviceps Harold, 1862

Copris laevigatus Gillet, 1927

Copris laiiformis Nguyen-phung & Cambefort, 1987

Copris laioides Boucomont, 1932

Copris laius Harold, 1868

Copris lannathai Hanboonsong & Masumoto & Ochi, 2003

Copris lecontei Matthews, 1962

Copris lugubris Boheman, 1858

Copris lunaris Linnaeus, 1758

Copris macacus Lansberge, 1886

Copris macclevei Warner, 1990

Copris macer Péringuey, 1901

Copris maesi Ratcliffe, 1998

Copris magicus Harold, 1881

Copris manni Ochi, Kon & Bai, 2009

Copris marcus Gillet, 1921

Copris martinae NGUYEN-PHUNG, 1988

Copris martinezi MATTHEWS & HALFFTER, 1968

Copris matthewsi Delgado & Kohlmann, 2001

Copris megasoma MATTHEWS & HALFFTER, 1959

Copris mesacanthus Harold, 1878

Copris mexicanus MATTHEWS & HALFFTER, 1959

Copris mindorensis Ochi, 1992

Copris minutissimus Kryzhanovskii & Medvedev, 1966

Copris minutus Drury, 1773

Copris misellus Péringuey, 1901

Copris miyakei Ochi & Kon, 2004

Copris moechus Leconte, 1854

Copris moffartsi Gillet, 1907

Copris mongkhoni Hanboonsong & Masumoto & Ochi, 2003

Copris montivagus Gillet, 1908

Copris morphaeus Gillet, 1932

Copris mourgliai Zelenka, 1993

Copris muticus Boheman, 1857

Copris neglectus Moxey, 1963

Copris nepos Gillet, 1908

Copris nevinsoni Waterhouse, 1891

Copris novaki Zelenka, 1992

Copris nubilosus Kohlmann & Cano & Delgado, 2003

Copris numa Lansberge, 1886

Copris obenbergeri Balthasar, 1933

Copris obesus Boheman, 1857

Copris ochus Motschulsky, 1860

Copris orion Klug, 1835

Copris orphanus Guérin-Méneville, 1847

Copris parapecuarius Ochi, Kon & Kawahara, 2008

Copris pauliani Nguyen-phung & Cambefort, 1986

Copris pecuarius Lewis, 1884

Copris pedarioides Lansberge, 1886

Copris phungae Cambefort, 1992

Copris phylax Gillet, 1908

Copris poggii Ochi & Kon, 2005

Copris potanini Semenov, 1891

Copris prevosti Josso, 2011

Copris propinquus Felsche, 1910

Copris pseudobootes Ferreira, 1962

Copris pseudochus Ochi & Kon, 2004

Copris pseudomoffartsi Moretto, 2011

Copris pseudosinicus Balthasar, 1958

Copris pueli MOLLANDIN DE BOISSY, 1905

Copris punctatus Gillet, 1910

Copris puncticollis Boheman, 1857

Copris punctipennis Boucomont, 1914

Copris punctiventris Waterhouse, 1891

Copris punctulatus Wiedemann, 1823

Copris punjabensis Gillet, 1921

Copris quasilaevigatus Ochi, Kon & Bai, 2007

Copris ramosiceps Gillet, 1921

Copris rebouchei Harold, 1869

Copris reflexus Fabricius, 1787

Copris remotus Leconte, 1866

Copris renwarti Nguyen-phung & Cambefort, 1986

Copris repertus Walker, 1858

Copris ribbei Lansberge, 1886

Copris riekoae Ochi & Kon, 2003

Copris ritsemae Harold, 1875

Copris ruandanus Balthasar, 1961

Copris rugosus Gillet, 1908

Copris ruricola Balthasar, 1933

Copris sabinus Gillet, 1910

Copris sacontala Redtenbacher, 1848

Copris sallei Harold, 1869

Copris sarpedon Harold, 1868

Copris saundersi Harold, 1869

Copris schoolmeestersi Ochi & Kon, 2010

Copris scorpio Zelenka, 1993

Copris serius NGUYEN-PHUNG, 1987

Copris sexdentatus Thunberg, 1818

Copris siamensis Gillet, 1910

Copris siangensis Biswas, 1980

Copris sierrensis Matthews, 1961

Copris signatus Walker, 1858

Copris simonettai Cambefort, 1992

Copris simplex Harold, 1869

Copris simulator Nguyen-phung & Cambefort, 1986

Copris singularis Nguyen-phung & Cambefort, 1986

Copris sinicus Hope, 1842

Copris sinon Olivier, 1789

Copris sodalis Walker, 1858

Copris sorex Balthasar, 1942

Copris sphaeropterus Harold, 1877

Copris spinator Harold, 1881

Copris subdolus Balthasar, 1958

Copris subpunctatus Gillet, 1910

Copris subsidens Péringuey, 1901

Copris sumatrensis Gillet, 1921

Copris surdus Arrow, 1931

Copris szechouanicus Balthasar, 1958

Copris tetraodon Gillet, 1910

Copris tridens Felsche, 1901

Copris tridentatus Solis & Kohlmann, 2003

Copris tripartitus Waterhouse, 1875

Copris truncatus Felsche, 1901

Copris tsukamotoi Ochi, Kon & Kawahara, 2008

Copris typhoeus Gerstaecker, 1884

Copris uenoi Hanboonsong & Masumoto & Ochi, 2003

Copris umbilicatus ABEILLE DE PERRIN, 1901

Copris urus Boheman, 1857

Copris usambaricus Gillet, 1908

Copris vankhaii NGUYEN-PHUNG, 1988

Copris victorini Boheman, 1857

Copris vietnamicus Kabakov, 1994

Copris vilhenai Ferreira, 1962

Copris vitalisi Gillet, 1921

Copris vrydaghi Ferreira, 1962

Copris warneri Mccleve & Kohlmann, 2005

Copris wiesei Kolbe, 1914

Copris yangi Ochi, Kon & Bai, 2007

Copris youngai Balthasar, 1967

Copris zhangi Ochi, Kon & Bai, 2009