Род Copris

Copris acutidens Motschulsky, 1860 [вид]

Copris afgoi Cambefort, 1992 [вид]

Copris agnus Sharp, 1875 [вид]

Copris algol NGUYEN-PHUNG, 1989 [вид]

Copris amabilis Kolbe, 1914 [вид]

Copris amyntor Klug, 1855 [вид]

Copris anceus Olivier, 1789 [вид]

Copris andrewesi Waterhouse, 1891 [вид]

Copris angusticornis Arrow, 1933 [вид]

Copris angustus Nguyen-phung & Cambefort, 1986 [вид]

Copris anomiopseoides Boucomont, 1924 [вид]

Copris antares Ferreira, 1958 [вид]

Copris apicepunctatus Balthasar, 1942 [вид]

Copris arcturus Gillet, 1907 [вид]

Copris ares Zelenka, 1993 [вид]

Copris arimotoi OCHI & ARAYA, 1992 [вид]

Copris arizonensis Schaeffer, 1906 [вид]

Copris armatus Harold, 1869 [вид]

Copris armeniacus Faldermann, 1835 [вид]

Copris armiger Gillet, 1910 [вид]

Copris arrowi Felsche, 1910 [вид]

Copris aspericollis Gillet, 1910 [вид]

Copris atropolitus Gillet, 1910 [вид]

Copris barclayi Ochi, Kon & Bai, 2009 [вид]

Copris basilewskyi Ferreira, 1962 [вид]

Copris basipunctatus Balthasar, 1942 [вид]

Copris bellator Chevrolat, 1844 [вид]

Copris bengalensis Gillet, 1911 [вид]

Copris bidens Kolbe, 1893 [вид]

Copris bihamatus Balthasar, 1965 [вид]

Copris bootes Klug, 1855 [вид]

Copris boucardi Harold, 1869 [вид]

Copris bovinus Gillet, 1908 [вид]

Copris brachypterus Nomura, 1964 [вид]

Copris brittoni Ferreira, 1962 [вид]

Copris caelatus Fabricius, 1794 [вид]

Copris caliginosus Kohlmann, Cano & Delgado, 2003 [вид]

Copris cambeforti NGUYEN-PHUNG, 1988 [вид]

Copris cambodiensis Ochi, Kon & Kawahara, 2008 [вид]

C. c. masumotoi

Copris camerunus Felsche, 1904 [вид]

Copris capensis Waterhouse, 1891 [вид]

Copris cariniceps Felsche, 1910 [вид]

Copris carinicus Gillet, 1910 [вид]

Copris carmelita Fabricius, 1801 [вид]

Copris cassius Péringuey, 1901 [вид]

Copris celebensis Lansberge, 1886 [вид]

Copris cheni Ochi, Kon & Bai, 2007 [вид]

Copris colmanti Gillet, 1908 [вид]

Copris complexus NGUYEN-PHUNG, 1989 [вид]

Copris compressipennis Gillet, 1910 [вид]

Copris confucius Harold, 1877 [вид]

Copris coriarius Gillet, 1907 [вид]

Copris cornifrons Boheman, 1860 [вид]

Copris corniger Sahlberg, 1823 [вид]

Copris corpulentus Gillet, 1910 [вид]

Copris costaricensis Gahan, 1894 [вид]

C. c. dolichocerus

Copris cribratus Gillet, 1927 [вид]

Copris cribricollis Gillet, 1910 [вид]

Copris curlettii Moretto, 2011 [вид]

Copris davisi Nguyen-phung & Cambefort, 1986 [вид]

Copris davisoni Waterhouse, 1891 [вид]

Copris decellei Cambefort, 1992 [вид]

Copris delicatus Arrow, 1931 [вид]

Copris denticulatus NGUYEN-PHUNG, 1988 [вид]

Copris diversus Waterhouse, 1891 [вид]

Copris doriae Harold, 1877 [вид]

Copris draco Arrow, 1906 [вид]

Copris dracunculus Fereira, 1959 [вид]

Copris dudleyi Cambefort, 1992 [вид]

Copris eburneus Cambefort, 1992 [вид]

Copris elpheneroides Felsche, 1910 [вид]

Copris elphenor Klug, 1855 [вид]

Copris erratus Lansberge, 1886 [вид]

C. e. masayukii

Copris evanidus Klug, 1855 [вид]

Copris excisus Waterhouse, 1891 [вид]

Copris fallaciosus Gillet, 1907 [вид]

Copris fallax Felsche, 1910 [вид]

Copris felschei Reitter, 1893 [вид]

Copris fidius Olivier, 1789 [вид]

Copris frankenbergerianus Balthasar, 1958 [вид]

Copris freyi Ferreira, 1961 [вид]

Copris fricator Fabricius, 1787 [вид]

C. f. cartwrighti

Copris fujiokai Ochi & Kon, 1996 [вид]

C. f. poringensis

Copris fukiensis Balthasar, 1952 [вид]

Copris furciceps Felsche, 1910 [вид]

Copris garambae Cambefort, 1992 [вид]

Copris gazellarum Gillet, 1918 [вид]

Copris gibbulus Lansberge, 1886 [вид]

C. g. borneensis

Copris gilleti Kolbe, 1907 [вид]

Copris gladiator Arrow, 1933 [вид]

Copris gopheri Hubbard, 1894 [вид]

Copris gracilis Waterhouse, 1891 [вид]

Copris graueri Kolbe, 1914 [вид]

Copris halffteri Matthews, 1959 [вид]

Copris harrisi Waterhouse, 1891 [вид]

Copris hartli Frey, 1975 [вид]

Copris hatayamai OCHI & ARAYA, 1992 [вид]

Copris hidakai Ochi & Kon, 1996 [вид]

Copris hispanus (Linnaeus, 1764) [вид]

C. h. cavolinii, C. h. hispanus, C. h. persianus

Copris howdeni MATTHEWS & HALFFTER, 1959 [вид]

Copris humilis Gillet, 1908 [вид]

Copris hybridus Gillet, 1910 [вид]

Copris igualensis Warner, 1990 [вид]

Copris illotus Balthasar, 1942 [вид]

Copris imitans Felsche, 1910 [вид]

Copris inaequabilis Zhang, 1997 [вид]

Copris incertus Say, 1835 [вид]

Copris inemarginatus Blatchley, 1918 [вид]

Copris inhalatus Quedenfeldt, 1884 [вид]

C. i. grandis, C. i. ngotto, C. i. perturbator, C. i. sanctaeluciae

Copris insidiosus Péringuey, 1901 [вид]

Copris integer Reiche, 1847 [вид]

Copris interioris Kolbe, 1897 [вид]

Copris iris Sharp, 1875 [вид]

Copris jacchoides Nguyen-phung & Cambefort, 1987 [вид]

Copris jacchus Fabricius, 1775 [вид]

Copris jahi Nguyen-phung & Cambefort, 1986 [вид]

Copris jucundus Gillet, 1932 [вид]

Copris kachinensis Ochi, Kon & Kawahara, 2008 [вид]

Copris kasagii Ochi & Kon, 1996 [вид]

Copris katangae Cambefort & Nguyen-phung, 1996 [вид]

Copris keralensis Gill, 1986 [вид]

Copris kiuchii Masumoto, 1989 [вид]

Copris klapperichi Balthasar, 1942 [вид]

Copris klugi Harold, 1869 [вид]

C. k. sierrensis

Copris kusakabei Ochi, Kon & Kawahara, 2005 [вид]

Copris laeviceps Harold, 1862 [вид]

Copris laevigatus Gillet, 1927 [вид]

Copris laiiformis Nguyen-phung & Cambefort, 1987 [вид]

Copris laioides Boucomont, 1932 [вид]

Copris laius Harold, 1868 [вид]

C. l. angusticornis

Copris lannathai Hanboonsong & Masumoto & Ochi, 2003 [вид]

Copris lecontei Matthews, 1962 [вид]

C. l. isthmiensis, C. l. lecontei

Copris lugubris Boheman, 1858 [вид]

Copris lunaris Linnaeus, 1758 [вид]

Copris macacus Lansberge, 1886 [вид]

Copris macclevei Warner, 1990 [вид]

Copris macer Péringuey, 1901 [вид]

Copris maesi Ratcliffe, 1998 [вид]

Copris magicus Harold, 1881 [вид]

Copris manni Ochi, Kon & Bai, 2009 [вид]

Copris marcus Gillet, 1921 [вид]

Copris martinae NGUYEN-PHUNG, 1988 [вид]

C. m. ntchisi

Copris martinezi MATTHEWS & HALFFTER, 1968 [вид]

Copris matthewsi Delgado & Kohlmann, 2001 [вид]

C. m. pacificus

Copris megaceratoides Waterhouse, 1891 [вид]

Copris megasoma MATTHEWS & HALFFTER, 1959 [вид]

Copris mesacanthus Harold, 1878 [вид]

C. m. transvaalensis

Copris mexicanus MATTHEWS & HALFFTER, 1959 [вид]

Copris mindorensis Ochi, 1992 [вид]

Copris minutissimus Kryzhanovskii & Medvedev, 1966 [вид]

Copris minutus Drury, 1773 [вид]

Copris misellus Péringuey, 1901 [вид]

Copris miyakei Ochi & Kon, 2004 [вид]

Copris moechus Leconte, 1854 [вид]

Copris moffartsi Gillet, 1907 [вид]

Copris mongkhoni Hanboonsong & Masumoto & Ochi, 2003 [вид]

Copris montivagus Gillet, 1908 [вид]

Copris morphaeus Gillet, 1932 [вид]

Copris mourgliai Zelenka, 1993 [вид]

Copris muticus Boheman, 1857 [вид]

Copris neglectus Moxey, 1963 [вид]

Copris nepos Gillet, 1908 [вид]

Copris nevinsoni Waterhouse, 1891 [вид]

Copris novaki Zelenka, 1992 [вид]

Copris nubilosus Kohlmann & Cano & Delgado, 2003 [вид]

Copris numa Lansberge, 1886 [вид]

Copris obenbergeri Balthasar, 1933 [вид]

Copris obesus Boheman, 1857 [вид]

Copris ochus Motschulsky, 1860 [вид]

Copris orion Klug, 1835 [вид]

C. o. caffer, C. o. centralis, C. o. overlaeti

Copris orphanus Guérin-Méneville, 1847 [вид]

Copris parapecuarius Ochi, Kon & Kawahara, 2008 [вид]

Copris pauliani Nguyen-phung & Cambefort, 1986 [вид]

Copris pecuarius Lewis, 1884 [вид]

C. p. continentalis

Copris pedarioides Lansberge, 1886 [вид]

Copris phungae Cambefort, 1992 [вид]

C. p. gabonicus

Copris phylax Gillet, 1908 [вид]

Copris poggii Ochi & Kon, 2005 [вид]

Copris potanini Semenov, 1891 [вид]

Copris prevosti Josso, 2011 [вид]

Copris propinquus Felsche, 1910 [вид]

Copris pseudobootes Ferreira, 1962 [вид]

Copris pseudochus Ochi & Kon, 2004 [вид]

Copris pseudomoffartsi Moretto, 2011 [вид]

Copris pseudosinicus Balthasar, 1958 [вид]

Copris pueli MOLLANDIN DE BOISSY, 1905 [вид]

Copris punctatus Gillet, 1910 [вид]

C. p. formosanus

Copris puncticollis Boheman, 1857 [вид]

Copris punctipennis Boucomont, 1914 [вид]

Copris punctiventris Waterhouse, 1891 [вид]

Copris punctulatus Wiedemann, 1823 [вид]

Copris punjabensis Gillet, 1921 [вид]

Copris quasilaevigatus Ochi, Kon & Bai, 2007 [вид]

Copris ramosiceps Gillet, 1921 [вид]

Copris rebouchei Harold, 1869 [вид]

Copris reflexus Fabricius, 1787 [вид]

Copris remotus Leconte, 1866 [вид]

C. r. dicyrtus, C. r. remotus

Copris renwarti Nguyen-phung & Cambefort, 1986 [вид]

Copris repertus Walker, 1858 [вид]

Copris ribbei Lansberge, 1886 [вид]

Copris riekoae Ochi & Kon, 2003 [вид]

Copris ritsemae Harold, 1875 [вид]

Copris ruandanus Balthasar, 1961 [вид]

Copris rugosus Gillet, 1908 [вид]

Copris ruricola Balthasar, 1933 [вид]

Copris sabinus Gillet, 1910 [вид]

Copris sacontala Redtenbacher, 1848 [вид]

Copris sallei Harold, 1869 [вид]

Copris sarpedon Harold, 1868 [вид]

Copris saundersi Harold, 1869 [вид]

Copris schoolmeestersi Ochi & Kon, 2010 [вид]

Copris scorpio Zelenka, 1993 [вид]

Copris serius NGUYEN-PHUNG, 1987 [вид]

Copris sexdentatus Thunberg, 1818 [вид]

Copris siamensis Gillet, 1910 [вид]

Copris siangensis Biswas, 1980 [вид]

Copris sierrensis Matthews, 1961 [вид]

Copris signatus Walker, 1858 [вид]

Copris simonettai Cambefort, 1992 [вид]

Copris simplex Harold, 1869 [вид]

Copris simulator Nguyen-phung & Cambefort, 1986 [вид]

Copris singularis Nguyen-phung & Cambefort, 1986 [вид]

Copris sinicus Hope, 1842 [вид]

Copris sinon Olivier, 1789 [вид]

Copris sodalis Walker, 1858 [вид]

Copris sorex Balthasar, 1942 [вид]

Copris sphaeropterus Harold, 1877 [вид]

Copris spinator Harold, 1881 [вид]

Copris subdolus Balthasar, 1958 [вид]

Copris subpunctatus Gillet, 1910 [вид]

Copris subsidens Péringuey, 1901 [вид]

Copris sumatrensis Gillet, 1921 [вид]

Copris surdus Arrow, 1931 [вид]

Copris szechouanicus Balthasar, 1958 [вид]

Copris tetraodon Gillet, 1910 [вид]

Copris tridens Felsche, 1901 [вид]

Copris tridentatus Solis & Kohlmann, 2003 [вид]

Copris tripartitus Waterhouse, 1875 [вид]

Copris truncatus Felsche, 1901 [вид]

Copris tsukamotoi Ochi, Kon & Kawahara, 2008 [вид]

Copris typhoeus Gerstaecker, 1884 [вид]

Copris uenoi Hanboonsong & Masumoto & Ochi, 2003 [вид]

Copris umbilicatus ABEILLE DE PERRIN, 1901 [вид]

Copris urus Boheman, 1857 [вид]

Copris usambaricus Gillet, 1908 [вид]

Copris vankhaii NGUYEN-PHUNG, 1988 [вид]

C. v. genopunctatus

Copris victorini Boheman, 1857 [вид]

Copris vietnamicus Kabakov, 1994 [вид]

Copris vilhenai Ferreira, 1962 [вид]

Copris vitalisi Gillet, 1921 [вид]

Copris vrydaghi Ferreira, 1962 [вид]

Copris warneri Mccleve & Kohlmann, 2005 [вид]

Copris wiesei Kolbe, 1914 [вид]

Copris yangi Ochi, Kon & Bai, 2007 [вид]

Copris youngai Balthasar, 1967 [вид]

Copris zhangi Ochi, Kon & Bai, 2009 [вид]