Genus Onthophagus
Insecta
urn:lsid:insecta.pro:taxonomy:830341
Daughter taxa
Amphionthophagus Martin-Piera & Zunino, 1983 [subgenus]
Furconthophagus Zunino, 1979 [subgenus]
Onthophagus Latreille, 1802 [subgenus]
Onthophagus aciculatulus, Onthophagus alluvius, Onthophagus antivagans, Onthophagus arnetti, Onthophagus batesi, Onthophagus brevifrons, Onthophagus browni, Onthophagus canelasensis, Onthophagus cartwrighti, Onthophagus cavernicollis, Onthophagus chevrolati, Onthophagus coahuilae, Onthophagus cochisus, Onthophagus concinnus, Onthophagus coproides, Onthophagus cribricollis, Onthophagus cuevensis, Onthophagus cyanellus, Onthophagus cynomysi, Onthophagus depressus, Onthophagus fuscus, Onthophagus hecate, Onthophagus hidalgus, Onthophagus hoepfneri, Onthophagus incensus, Onthophagus knausi, Onthophagus knulli, Onthophagus landolti, Onthophagus lecontei, Onthophagus mcclevei, Onthophagus medorensis, Onthophagus mextexus, Onthophagus oklahomensis, Onthophagus orpheus, Onthophagus pennsylvanicus, Onthophagus polyphemi, Onthophagus pseudofuscus, Onthophagus schaefferi, Onthophagus striatulus, Onthophagus subaeneus, Onthophagus subopacus, Onthophagus subtropicus, Onthophagus taurus, Onthophagus tuberculifrons, Onthophagus velutinus
Relictonthophagus Kabakov, 1979 [subgenus]
Trichonthophagus Zunino, 1979 [subgenus]
Onthophagus abacus Boucomont, 1921 [species]
Onthophagus abas Balthasar, 1946 [species]
Onthophagus abeillei D'orbigny, 1897 [species]
Onthophagus abreui Arrow, 1931 [species]
Onthophagus abruptus D'orbigny, 1913 [species]
Onthophagus absyrtus Balthasar, 1946 [species]
Onthophagus abyssinicus Gillet, 1925 [species]
Onthophagus academus Balthasar, 1946 [species]
Onthophagus accedens Lansberge, 1883 [species]
Onthophagus acerus Gillet, 1930 [species]
Onthophagus aciculatus Fahraeus, 1857 [species]
Onthophagus acrisius Balthasar, 1946 [species]
Onthophagus acuminatus Harold, 1880 [species]
Onthophagus acuticornis Endrödi, 1973 [species]
Onthophagus acutus D'orbigny, 1908 [species]
Onthophagus adelaidae Hope, 1846 [species]
Onthophagus adelphus Gillet, 1930 [species]
Onthophagus admetus Balthasar, 1946 [species]
Onthophagus adornatus D'orbigny, 1904 [species]
Onthophagus adspersus D'orbigny, 1908 [species]
Onthophagus aegrotus Balthasar, 1967 [species]
Onthophagus aemulus Gillet, 1930 [species]
Onthophagus aeneoniger D'orbigny, 1913 [species]
Onthophagus aeneopiceus D'orbigny, 1902 [species]
Onthophagus aenescens Wiedemann, 1823 [species]
Onthophagus aequatus Péringuey, 1901 [species]
Onthophagus aequepubens D'orbigny, 1905 [species]
Onthophagus aerarius Reitter, 1892 [species]
Onthophagus aereidorsis D'orbigny, 1902 [species]
Onthophagus aeremicans D'orbigny, 1904 [species]
Onthophagus aereomaculatus Boucomont, 1914 [species]
Onthophagus aerestriatus D'orbigny, 1913 [species]
Onthophagus aeruginosus Roth, 1851 [species]
Onthophagus aerumnosus Balthasar, 1963 [species]
Onthophagus aesopus Lansberge, 1882 [species]
Onthophagus aethiopicus D'orbigny, 1902 [species]
Onthophagus affinis Gillet, 1930 [species]
Onthophagus afghanus Petrovitz, 1961 [species]
Onthophagus africanus Lansberge, 1886 [species]
Onthophagus agaricophilus Arrow, 1931 [species]
Onthophagus agnus Gillet, 1925 [species]
Onthophagus ahenicollis D'orbigny, 1902 [species]
Onthophagus ahenomicans D'orbigny, 1902 [species]
Onthophagus akhaus Masumoto, Ochi & Hanboonsong, 2007 [species]
Onthophagus akinini Koenig, 1889 [species]
Onthophagus albarracinus Baraud, 1979 [species]
Onthophagus albicapillus D'Orbigny, 1902 [species]
Onthophagus albicomus D'orbigny, 1908 [species]
Onthophagus albicornis Palisot de Beauvois, 1805 [species]
Onthophagus albipennis Péringuey, 1908 [species]
Onthophagus albipodex D'orbigny, 1902 [species]
Onthophagus alcedo d'Orbigny, 1913 [species]
Onthophagus alcyon Klug, 1855 [species]
Onthophagus alcyonides d'Orbigny, 1913 [species]
Onthophagus alexeevi Tarasov, Krikken & Huijbregts, 2010 [species]
Onthophagus alfuricus Huijbregts & Krikken, 2012 [species]
Onthophagus alienus Frey, 1958 [species]
Onthophagus allojavanus Huijbregts & Krikken, 2011 [species]
Onthophagus alluaudi D'orbigny, 1902 [species]
Onthophagus aloysiellus Zunino, 1977 [species]
Onthophagus alquirta Matthews, 1972 [species]
Onthophagus alternans Raffray, 1877 [species]
Onthophagus altidorsis D'orbigny, 1905 [species]
Onthophagus altilamina D'orbigny, 1905 [species]
Onthophagus altivagans Howden & Génier, 2004 [species]
Onthophagus amamiensis Nomura, 1965 [species]
Onthophagus amblychromatus Marcus, 1917 [species]
Onthophagus amicus Gillet, 1925 [species]
Onthophagus amirus Kabakov, 1982 [species]
Onthophagus amoenus D'orbigny, 1908 [species]
Onthophagus amphicoma Boucomont, 1914 [species]
Onthophagus amphinasus Arrow, 1931 [species]
Onthophagus amphioxus Arrow, 1933 [species]
Onthophagus amplexus Sharp, 1875 [species]
Onthophagus amplipennis D'orbigny, 1905 [species]
Onthophagus anatolicus Petrovitz, 1962 [species]
Onthophagus anceyi Boucomont, 1921 [species]
Onthophagus anchommatus Lea, 1923 [species]
Onthophagus andersoni HOWDEN & GILL, 1987 [species]
Onthophagus andonarensis Boucomont, 1914 [species]
Onthophagus andrewdavisi Huijbregts & Krikken, 2009 [species]
Onthophagus andrewesi Arrow, 1931 [species]
Onthophagus androgynus D'orbigny, 1905 [species]
Onthophagus anewtoni Howden & Génier, 2004 [species]
Onthophagus angolensis Paulian, 1937 [species]
Onthophagus angorensis Petrovitz, 1963 [species]
Onthophagus anguicorius Boucomont, 1914 [species]
Onthophagus angularis D'orbigny, 1908 [species]
Onthophagus anguliceps Boucomont, 1914 [species]
Onthophagus angulicornis D'orbigny, 1908 [species]
Onthophagus angustatus Boucomont, 1914 [species]
Onthophagus anisocerus Erichson, 1842 [species]
Onthophagus anitidus Ochi & Kon, 2005 [species]
Onthophagus annoyeri Moretto, 2010 [species]
Onthophagus anogeissii Cambefort, 1984 [species]
Onthophagus anomalicollis Frey, 1973 [species]
Onthophagus anomalipes D'orbigny, 1902 [species]
Onthophagus antennalis Frey, 1961 [species]
Onthophagus anthracinus Harold, 1873 [species]
Onthophagus antillarum Arrow, 1903 [species]
Onthophagus antilocapra Balthasar, 1937 [species]
Onthophagus antoinei Walter, 1990 [species]
Onthophagus aokii Nomura, 1976 [species]
Onthophagus aper Sharp, 1875 [species]
Onthophagus aphodioides Lansberge, 1883 [species]
Onthophagus apicetinctus D'orbigny, 1898 [species]
Onthophagus apiciosus D'orbigny, 1902 [species]
Onthophagus apilularius Masumoto, 1995 [species]
Onthophagus apterus Matthews, 1972 [species]
Onthophagus apunneea Masumoto, Ochi & Hanboonsong, 2007 [species]
Onthophagus arai Masumoto, 1989 [species]
Onthophagus arayai Ochi & Kon, 2007 [species]
Onthophagus arboreus Arrow, 1931 [species]
Onthophagus arcifer D'orbigny, 1904 [species]
Onthophagus ardoini Frey, 1971 [species]
Onthophagus areolatus D'orbigny, 1905 [species]
Onthophagus argyropygus Gillet, 1927 [species]
Onthophagus aries Kirby, 1825 [species]
Onthophagus arkoola STOREY & WEIR, 1990 [species]
Onthophagus armatus Blanchard, 1853 [species]
Onthophagus armicollis D'Orbigny, 1904 [species]
Onthophagus arnoldii Kabakov, 1982 [species]
Onthophagus arrilla Matthews, 1972 [species]
Onthophagus arunachalensis Biswas & Chatterjee, 1985 [species]
Onthophagus arunensis Scheuern, 1995 [species]
Onthophagus aschenborni Frey, 1975 [species]
Onthophagus ashanticola Frey, 1973 [species]
Onthophagus asiaticus Endrödi, 1973 [species]
Onthophagus asimilis Péringuey, 1901 [species]
Onthophagus asper Macleay, 1864 [species]
Onthophagus asperatus D'orbigny, 1905 [species]
Onthophagus aspericeps D'orbigny, 1908 [species]
Onthophagus aspericollis Lansberge, 1885 [species]
Onthophagus asperipennis D'orbigny, 1902 [species]
Onthophagus aspernatus D'orbigny, 1907 [species]
Onthophagus asperodorsatus HOWDEN & GILL, 1993 [species]
Onthophagus asperrimus D'orbigny, 1902 [species]
Onthophagus asperulus D'orbigny, 1905 [species]
Onthophagus astigma D'orbigny, 1904 [species]
Onthophagus ater Waterhouse, 1875 [species]
Onthophagus athiensis D'orbigny, 1913 [species]
Onthophagus atratus Kabakov, 1983 [species]
Onthophagus atricapillus D'Orbigny, 1908 [species]
Onthophagus atriclaviger d'Orbigny, 1905 [species]
Onthophagus atricolor D'orbigny, 1907 [species]
Onthophagus atridorsis D'orbigny, 1902 [species]
Onthophagus atriglabrus HOWDEN & GILL, 1987 [species]
Onthophagus atripennis Waterhouse, 1875 [species]
Onthophagus atroaereus D'orbigny, 1908 [species]
Onthophagus atrofasciatus D'orbigny, 1905 [species]
Onthophagus atronitidus D'orbigny, 1902 [species]
Onthophagus atropolitus D'Orbigny, 1902 [species]
Onthophagus atrosericeus Boucomont, 1932 [species]
Onthophagus atrosetosus D'Orbigny, 1902 [species]
Onthophagus atrostriatus D'orbigny, 1913 [species]
Onthophagus atrovirens D'orbigny, 1905 [species]
Onthophagus atrovittatus D'orbigny, 1908 [species]
Onthophagus atrox Harold, 1867 [species]
Onthophagus auratus Fabricius, 1801 [species]
Onthophagus aureiceps d'Orbigny, 1902 [species]
Onthophagus aureofuscus Bates, 1887 [species]
Onthophagus aureopilosus Boucomont, 1914 [species]
Onthophagus auriculatus Klug, 1855 [species]
Onthophagus aurifex Harold, 1877 [species]
Onthophagus auritus Erichson, 1842 [species]
Onthophagus australis Guérin-Méneville, 1830 [species]
Onthophagus avocetta Arrow, 1933 [species]
Onthophagus avocettoides Kabakov, 1983 [species]
Onthophagus axillaris Boheman, 1860 [species]
Onthophagus aztecus ZUNINO & HALFFTER, 1988 [species]
Onthophagus azusae Ochi & Kon, 2006 [species]
Onthophagus babaulti D'orbigny, 1915 [species]
Onthophagus babirussa Eschscholtz, 1822 [species]
Onthophagus badeni Harold, 1875 [species]
Onthophagus baenzigeri Masumoto, Ochi & Hanboonsong, 2007 [species]
Onthophagus baeri Boucomont, 1914 [species]
Onthophagus bakeri Boucomont, 1919 [species]
Onthophagus balawaicus Scheuern, 1995 [species]
Onthophagus baloghi Balthasar, 1967 [species]
Onthophagus balthasari Vsetecka, 1939 [species]
Onthophagus bambra Matthews, 1972 [species]
Onthophagus bandamai Cambefort, 1984 [species]
Onthophagus bangueyensis Boucomont, 1914 [species]
Onthophagus baoule Cambefort, 1984 [species]
Onthophagus baraudi Nicolas, 1964 [species]
Onthophagus barbieri Paulian, 1978 [species]
Onthophagus barretti Génier & Howden, 1999 [species]
Onthophagus barriorum Walter, 1990 [species]
Onthophagus bartosi Balthasar, 1966 [species]
Onthophagus basakata WALTER & CAMBEFORT, 1977 [species]
Onthophagus basicruentatus Goidanich, 1926 [species]
Onthophagus basilewskyi Frey, 1961 [species]
Onthophagus basipustulatus Heyden, 1889 [species]
Onthophagus bassariscus ZUNINO & HALFFTER, 1988 [species]
Onthophagus bateke WALTER & CAMBEFORT, 1977 [species]
Onthophagus batillifer Harold, 1875 [species]
Onthophagus batui Huijbregts & Krikken, 2009 [species]
Onthophagus bayeri Balthasar, 1942 [species]
Onthophagus baykanus Kabakov, 1994 [species]
Onthophagus bechynei Frey, 1953 [species]
Onthophagus beelarong STOREY & WEIR, 1990 [species]
Onthophagus beesoni Arrow, 1931 [species]
Onthophagus beiranus Péringuey, 1908 [species]
Onthophagus belinga Walter, 1989 [species]
Onthophagus bellus D'orbigny, 1905 [species]
Onthophagus belorhinus Bates, 1887 [species]
Onthophagus benedictorum WALTER & CAMBEFORT, 1977 [species]
Onthophagus bengalensis Harold, 1886 [species]
Onthophagus bengali Gordon & Oppenheimer, 1977 [species]
Onthophagus benguellianus Paulian, 1937 [species]
Onthophagus bennigseni Paulian, 1937 [species]
Onthophagus bequaerti D'orbigny, 1915 [species]
Onthophagus bergeri Frey, 1975 [species]
Onthophagus bernaudi Cambefort & nicolas, 1991 [species]
Onthophagus betschuanus Frey, 1975 [species]
Onthophagus biarmatus D'Orbigny, 1908 [species]
Onthophagus bicallifrons D'orbigny, 1902 [species]
Onthophagus bicallosus Klug, 1855 [species]
Onthophagus bicarinaticeps Lea, 1923 [species]
Onthophagus bicarinatus Gomes Alves, 1944 [species]
Onthophagus bicavicollis Lea, 1923 [species]
Onthophagus bicavifrons D'orbigny, 1902 [species]
Onthophagus bicolor Raffray, 1877 [species]
Onthophagus biconifer D'orbigny, 1905 [species]
Onthophagus bicornis Macleay, 1888 [species]
Onthophagus bicristatus D'orbigny, 1905 [species]
Onthophagus bicristiger D'orbigny, 1913 [species]
Onthophagus bicuneus Kabakov, 2008 [species]
Onthophagus bidens Olivier, 1789 [species]
Onthophagus bidentatus Drapiez, 1819 [species]
Onthophagus bidentifrons D'orbigny, 1902 [species]
Onthophagus biexcavatus D'orbigny, 1898 [species]
Onthophagus bifasciatus Fabricius, 1781 [species]
Onthophagus bifidicornis D'orbigny, 1902 [species]
Onthophagus bifidus Reiche, 1849 [species]
Onthophagus bifrons D'orbigny, 1905 [species]
Onthophagus bilingula Balthasar, 1965 [species]
Onthophagus bimarginatus D'orbigny, 1902 [species]
Onthophagus bimetallicus D'orbigny, 1907 [species]
Onthophagus bindaree STOREY & WEIR, 1990 [species]
Onthophagus binodis Thunberg, 1818 [species]
Onthophagus binodosus D'orbigny, 1908 [species]
Onthophagus binodulus D'orbigny, 1913 [species]
Onthophagus binyana STOREY & WEIR, 1990 [species]
Onthophagus biplagiatus Thomson, 1858 [species]
Onthophagus birugatus D'orbigny, 1902 [species]
Onthophagus birugifer D'orbigny, 1908 [species]
Onthophagus bisbicornis D'orbigny, 1909 [species]
Onthophagus biscarinulatus Huijbregts & Krikken, 2012 [species]
Onthophagus bisectus Arrow, 1931 [species]
Onthophagus bisignatus D'orbigny, 1913 [species]
Onthophagus bison Boucomont, 1919 [species]
Onthophagus bistiniocelloides Krikken, 1986 [species]
Onthophagus bituber D'orbigny, 1904 [species]
Onthophagus bituberans D'orbigny, 1905 [species]
Onthophagus bituberculatus Olivier, 1789 [species]
Onthophagus bivertex Heyden, 1887 [species]
Onthophagus blackburni Shipp, 1895 [species]
Onthophagus blackwoodensis Blackburn, 1891 [species]
Onthophagus blanchardi Harold, 1869 [species]
Onthophagus blumei Lansberge, 1883 [species]
Onthophagus bocandei D'orbigny, 1904 [species]
Onthophagus bokiaunus Masumoto, 1995 [species]
Onthophagus bomberaianus Balthasar, 1969 [species]
Onthophagus bonengus Kabakov, 1994 [species]
Onthophagus bonorae Zunino, 1976 [species]
Onthophagus bonsae Zunino, 1976 [species]
Onthophagus boops D'orbigny, 1905 [species]
Onthophagus boranus Gestro, 1895 [species]
Onthophagus borassi Cambefort, 1984 [species]
Onthophagus borneensis Harold, 1877 [species]
Onthophagus bornemisszai Matthews, 1972 [species]
Onthophagus bornemisszanus Matthews, 1972 [species]
Onthophagus bottegi Gestro, 1895 [species]
Onthophagus boucomonti Paulian, 1931 [species]
Onthophagus bourgognei Paulian, 1945 [species]
Onthophagus bovinus Péringuey, 1892 [species]
Onthophagus brachypterus ZUNINO & HALFFTER, 1997 [species]
Onthophagus brazzavillianus Balthasar, 1974 [species]
Onthophagus brendelli Ochi, Kon & Barclay, 2008 [species]
Onthophagus breviceps D'orbigny, 1902 [species]
Onthophagus brevicollis Arrow, 1907 [species]
Onthophagus breviconus Génier & Howden, 1999 [species]
Onthophagus brevigena D'orbigny, 1902 [species]
Onthophagus brevipennis D'orbigny, 1902 [species]
Onthophagus brevisetis D'orbigny, 1915 [species]
Onthophagus brittoni Paulian, 1948 [species]
Onthophagus brivioi Frey, 1973 [species]
Onthophagus bronzeus Arrow, 1907 [species]
Onthophagus brooksi Matthews, 1972 [species]
Onthophagus brunellii Müller, 1942 [species]
Onthophagus brutus Arrow, 1931 [species]
Onthophagus bubalus Harold, 1867 [species]
Onthophagus buculus Mannerheim, 1829 [species]
Onthophagus bufulus Arrow, 1941 [species]
Onthophagus bunamin Matthews, 1972 [species]
Onthophagus bundara STOREY & WEIR, 1990 [species]
Onthophagus bundutuhanensis Ochi, Kon & Barclay, 2008 [species]
Onthophagus burchelli D'orbigny, 1908 [species]
Onthophagus busiris Balthasar, 1969 [species]
Onthophagus bytinskii Balthasar, 1960 [species]
Onthophagus caelator Balthasar, 1967 [species]
Onthophagus caesariatus Boucomont, 1921 [species]
Onthophagus caffrarius D'orbigny, 1902 [species]
Onthophagus calamophilus Krikken, 1977 [species]
Onthophagus calcaratus Boucomont, 1919 [species]
Onthophagus calchas Balthasar, 1967 [species]
Onthophagus calliger D'orbigny, 1913 [species]
Onthophagus callosipennis Boucomont, 1930 [species]
Onthophagus cambeforti Palestrini, 1992 [species]
Onthophagus cameloides D'Orbigny, 1898 [species]
Onthophagus camerunicus D'orbigny, 1905 [species]
Onthophagus cancer Lansberge, 1886 [species]
Onthophagus capella Kirby, 1818 [species]
Onthophagus capelliformis Gillet, 1925 [species]
Onthophagus capellinus Frey, 1963 [species]
Onthophagus capitatus Castelnau, 1840 [species]
Onthophagus capitosus Harold, 1867 [species]
Onthophagus caprai Frey, 1956 [species]
Onthophagus carayoni Cambefort, 1986 [species]
Onthophagus carcharias Harold, 1875 [species]
Onthophagus carinensis Boucomont, 1914 [species]
Onthophagus carinicollis Raffray, 1877 [species]
Onthophagus carinidorsis D'orbigny, 1908 [species]
Onthophagus carinifer D'orbigny, 1904 [species]
Onthophagus carinulatus Harold, 1877 [species]
Onthophagus carmodensis Blackburn, 1907 [species]
Onthophagus carpanetoi Pittino, 1982 [species]
Onthophagus carpophilus Pereira & Halffter, 1961 [species]
Onthophagus castetsi Lansberge, 1887 [species]
Onthophagus catenatus Lansberge, 1883 [species]
Onthophagus catharinensis Paulian, 1936 [species]
Onthophagus catta Fabricius, 1787 [species]
Onthophagus cavia Boucomont, 1914 [species]
Onthophagus caviceps Frey, 1962 [species]
Onthophagus cavidorsis d'Orbigny, 1902 [species]
Onthophagus cavifrons Harold, 1886 [species]
Onthophagus cavivertex D'orbigny, 1913 [species]
Onthophagus centricornis Fabricius, 1798 [species]
Onthophagus centurio Lansberge, 1885 [species]
Onthophagus cephalophi Cambefort, 1984 [species]
Onthophagus ceratotherium Janssens, 1953 [species]
Onthophagus cervenkai Kabakov, 2008 [species]
Onthophagus cervicapra Boucomont, 1914 [species]
Onthophagus cervus Fabricius, 1798 [species]
Onthophagus chaingraiensis Masumoto, Ochi & Hanboonsong, 2007 [species]
Onthophagus chaiyaphumensis Masumoto, Ochi & Hanboonsong, 2002 [species]
Onthophagus chalcostolus D'Orbigny, 1902 [species]
Onthophagus championi Bates, 1887 [species]
Onthophagus chandrai Ochi, 2007 [species]
Onthophagus changshouensis Zhang, 1997 [species]
Onthophagus chapaensis Kabakov, 1983 [species]
Onthophagus chebaicus Boucomont, 1919 [species]
Onthophagus cheesmanae Arrow, 1941 [species]
Onthophagus chepara Matthews, 1972 [species]
Onthophagus cheyi Ochi & Kon, 2006 [species]
Onthophagus chiapanecus ZUNINO & HALFFTER, 1988 [species]
Onthophagus chineicus Kabakov, 1998 [species]
Onthophagus chinensis Balthasar, 1952 [species]
Onthophagus chirindanus D'orbigny, 1913 [species]
Onthophagus chloroderus D'orbigny, 1905 [species]
Onthophagus chlorophanus D'orbigny, 1902 [species]
Onthophagus choanicus D'orbigny, 1904 [species]
Onthophagus chremes Balthasar, 1969 [species]
Onthophagus chryses Bates, 1887 [species]
Onthophagus chrysoderus D'orbigny, 1905 [species]
Onthophagus chrysopes Bates, 1888 [species]
Onthophagus chrysurus Arrow, 1931 [species]
Onthophagus chumphonensis Masumoto, Ochi & Hanboonsong, 2008 [species]
Onthophagus cicer Balthasar, 1964 [species]
Onthophagus cincticollis D'orbigny, 1902 [species]
Onthophagus cinctifrons D'orbigny, 1902 [species]
Onthophagus cinctipennis Quedenfeldt, 1884 [species]
Onthophagus cineraceus D'orbigny, 1902 [species]
Onthophagus circulator Reitter, 1891 [species]
Onthophagus circulifer Arrow, 1931 [species]
Onthophagus circumdatus D'orbigny, 1913 [species]
Onthophagus citreum Boucomont, 1919 [species]
Onthophagus civettae Cambefort, 1984 [species]
Onthophagus clavisetis D'orbigny, 1905 [species]
Onthophagus clermonti Paulian, 1931 [species]
Onthophagus clitellarius D'orbigny, 1908 [species]
Onthophagus clitellifer Reitter, 1894 [species]
Onthophagus clivimerus Huijbregts & Krikken, 2011 [species]
Onthophagus cludtsi Ochi, 2003 [species]
Onthophagus clusifrons D'orbigny, 1905 [species]
Onthophagus clypealis Lea, 1923 [species]
Onthophagus clypeatus Blanchard, 1846 [species]
Onthophagus coenobita Herbst, 1783 [species]
Onthophagus coeruleicollis Arrow, 1907 [species]
Onthophagus cognatus Boucomont, 1921 [species]
Onthophagus coiffaiti Frey, 1970 [species]
Onthophagus colasi Paulian, 1937 [species]
Onthophagus colffsi Lansberge, 1883 [species]
Onthophagus collaris Kabakov, 1983 [species]
Onthophagus collinsi Krikken & Huijbregts, 1987 [species]
Onthophagus colmanti D'Orbigny, 1908 [species]
Onthophagus columbianus Boucomont, 1932 [species]
Onthophagus comatulus D'orbigny, 1905 [species]
Onthophagus cometes Bates, 1888 [species]
Onthophagus commottoides Kabakov, 1998 [species]
Onthophagus comottoi Lansberge, 1885 [species]
Onthophagus compactus Arrow, 1933 [species]
Onthophagus comperei Blackburn, 1903 [species]
Onthophagus compositus Lea, 1923 [species]
Onthophagus compressus Guérin-Méneville, 1855 [species]
Onthophagus concavifrons D'orbigny, 1904 [species]
Onthophagus concolor Sharp, 1878 [species]
Onthophagus concursator Balthasar, 1960 [species]
Onthophagus confertus Péringuey, 1908 [species]
Onthophagus confluens D'orbigny, 1902 [species]
Onthophagus confossus D'Orbigny, 1915 [species]
Onthophagus conradsi Balthasar, 1937 [species]
Onthophagus consentaneus Harold, 1867 [species]
Onthophagus conspersus Reitter, 1893 [species]
Onthophagus conspicuus Macleay, 1864 [species]
Onthophagus contiguicornis D'orbigny, 1913 [species]
Onthophagus convexicollis Boheman, 1858 [species]
Onthophagus convexus D'orbigny, 1908 [species]
Onthophagus cooloola STOREY & WEIR, 1990 [species]
Onthophagus coomani Paulian, 1931 [species]
Onthophagus coorgensis Arrow, 1931 [species]
Onthophagus coprimorphoides Huijbregts & Krikken, 2012 [species]
Onthophagus coprimorphus Gillet, 1930 [species]
Onthophagus coptorhinodes Péringuey, 1901 [species]
Onthophagus coracinoides Kabakov, 1983 [species]
Onthophagus coracinus Boucomont, 1914 [species]
Onthophagus coriaceoumbrosus Kohlmann & Solis, 2001 [species]
Onthophagus corniculatus Reiche, 1847 [species]
Onthophagus corniculiger D'orbigny, 1913 [species]
Onthophagus cornifrons Thomson, 1858 [species]
Onthophagus cornutus Ferreira, 1971 [species]
Onthophagus coronatus D'orbigny, 1902 [species]
Onthophagus corrosus Bates, 1887 [species]
Onthophagus coscineus Bates, 1887 [species]
Onthophagus costatus D'orbigny, 1913 [species]
Onthophagus costifer Gillet, 1930 [species]
Onthophagus costiger D'orbigny, 1915 [species]
Onthophagus costilatus D'orbigny, 1908 [species]
Onthophagus costulicollis Frey, 1975 [species]
Onthophagus crantor Balthasar, 1967 [species]
Onthophagus crassicollis Boucomont, 1913 [species]
Onthophagus cratippus Balthasar, 1969 [species]
Onthophagus creber D'Orbigny, 1905 [species]
Onthophagus cribellum D'orbigny, 1902 [species]
Onthophagus cribratus Lansberge, 1883 [species]
Onthophagus cribripennis D'orbigny, 1902 [species]
Onthophagus criniger D'orbigny, 1904 [species]
Onthophagus cristatus D'orbigny, 1905 [species]
Onthophagus croesulus Bates, 1888 [species]
Onthophagus crotchi Harold, 1871 [species]
Onthophagus crucenotatus D'orbigny, 1905 [species]
Onthophagus cruciatus Ménétriès, 1832 [species]
Onthophagus cruciger Macleay, 1888 [species]
Onthophagus cruentatus Klug, 1855 [species]
Onthophagus cryptodicranius Kohlmann & Solis, 2001 [species]
Onthophagus cryptogenus Boucomont, 1914 [species]
Onthophagus cuneus Boucomont, 1924 [species]
Onthophagus cuniculus Macleay, 1864 [species]
Onthophagus cupreiceps Arrow, 1907 [species]
Onthophagus cupreopastillatus Ochi & Kon, 2006 [species]
Onthophagus cupreovirens D'orbigny, 1904 [species]
Onthophagus cupreus Harold, 1880 [species]
Onthophagus cupricollis Péringuey, 1888 [species]
Onthophagus curtipilis D'orbigny, 1905 [species]
Onthophagus curtulus Frey, 1961 [species]
Onthophagus curvicarinatus Boucomont, 1914 [species]
Onthophagus curvicornis Latreille, 1812 [species]
Onthophagus curvifrons D'orbigny, 1906 [species]
Onthophagus curvilamina Müller, 1942 [species]
Onthophagus curvispina Reitter, 1893 [species]
Onthophagus cuspidatus d'Orbigny, 1913 [species]
Onthophagus cyaneiceps D'orbigny, 1913 [species]
Onthophagus cyaneoniger D'orbigny, 1902 [species]
Onthophagus cyaneus Kabakov, 2006 [species]
Onthophagus cyanochlorus D'orbigny, 1902 [species]
Onthophagus cyanopterus D'orbigny, 1905 [species]
Onthophagus cyclographus Bates, 1887 [species]
Onthophagus cyobioides Boucomont, 1921 [species]
Onthophagus dacatrai Pittino, 2004 [species]
Onthophagus dama Fabricius, 1798 [species]
Onthophagus damaki Masumoto, Hanboonsong & Ochi, 2002 [species]
Onthophagus dandalu Matthews, 1972 [species]
Onthophagus danumcupreus Krikken & Huijbregts, 2009 [species]
Onthophagus danumensis Ochi, Kon & Barclay, 2009 [species]
Onthophagus danumicus Huijbregts & Krikken, 2012 [species]
Onthophagus dapcauensis Boucomont, 1921 [species]
Onthophagus dapitanensis Boucomont, 1919 [species]
Onthophagus darlingtoni Matthews, 1972 [species]
Onthophagus davisi Frey, 1975 [species]
Onthophagus dayacus Boucomont, 1914 [species]
Onthophagus debilis D'orbigny, 1905 [species]
Onthophagus deccanensis Frey, 1962 [species]
Onthophagus decedens Péringuey, 1904 [species]
Onthophagus decens Balthasar, 1964 [species]
Onthophagus declivicollis D'orbigny, 1902 [species]
Onthophagus declivis Harold, 1869 [species]
Onthophagus decolor D'orbigny, 1902 [species]
Onthophagus decoratus D'orbigny, 1905 [species]
Onthophagus decorsei D'orbigny, 1908 [species]
Onthophagus dedecor Wallengren, 1881 [species]
Onthophagus deflexicollis Lansberge, 1883 [species]
Onthophagus deflexus D'orbigny, 1908 [species]
Onthophagus dejongi Krikken, 1986 [species]
Onthophagus delahayei Josso, 2011 [species]
Onthophagus delicatulus Raffray, 1877 [species]
Onthophagus delicatus D'orbigny, 1915 [species]
Onthophagus dellacasai Pittino & Mariani, 1981 [species]
Onthophagus delphinensis D'orbigny, 1913 [species]
Onthophagus demarzi Frey, 1959 [species]
Onthophagus demeyeri Moretto, 2009 [species]
Onthophagus densatus Frey, 1963 [species]
Onthophagus densepunctatus Frey, 1963 [species]
Onthophagus densipilis D'orbigny, 1902 [species]
Onthophagus denticollis Lansberge, 1883 [species]
Onthophagus denticornis Boucomont, 1914 [species]
Onthophagus denticulatus D'orbigny, 1902 [species]
Onthophagus denudatus D'orbigny, 1902 [species]
Onthophagus depilatus D'orbigny, 1913 [species]
Onthophagus depilis D'orbigny, 1902 [species]
Onthophagus deplanatus Lansberge, 1883 [species]
Onthophagus depressicollis Boucomont, 1914 [species]
Onthophagus depressifrons D'orbigny, 1915 [species]
Onthophagus depressipennis Cambefort, 1975 [species]
Onthophagus derasus D'orbigny, 1902 [species]
Onthophagus desaegeri Frey, 1961 [species]
Onthophagus desectus Macleay, 1871 [species]
Onthophagus deterrens Péringuey, 1901 [species]
Onthophagus devagiriensis Schoolmeesters & Thomas, 2006 [species]
Onthophagus devexicornis Gillet, 1930 [species]
Onthophagus devexus Macleay, 1888 [species]
Onthophagus dhanjuricus Frey, 1973 [species]
Onthophagus diabolicus Harold, 1877 [species]
Onthophagus diadematus D'orbigny, 1902 [species]
Onthophagus dicax Balthasar, 1966 [species]
Onthophagus dicella Bates, 1888 [species]
Onthophagus dicranius Bates, 1887 [species]
Onthophagus dicranocerus Gillet, 1925 [species]
Onthophagus dicranoides Balthasar, 1939 [species]
Onthophagus difficilis Le Guillou, 1844 [species]
Onthophagus digitatus Arrow, 1931 [species]
Onthophagus dignus Gillet, 1930 [species]
Onthophagus dilutus D'orbigny, 1905 [species]
Onthophagus dinjeera Storey & Weir, 1990 [species]
Onthophagus dinoderus D'orbigny, 1913 [species]
Onthophagus discedens Sharp, 1875 [species]
Onthophagus discolor Hope, 1841 [species]
Onthophagus discovirens D'orbigny, 1902 [species]
Onthophagus discretus Péringuey, 1901 [species]
Onthophagus dispar Péringuey, 1901 [species]
Onthophagus dissentaneus Balthasar, 1964 [species]
Onthophagus distichus Roth, 1851 [species]
Onthophagus ditus Péringuey, 1901 [species]
Onthophagus diversiformis Boucomont, 1914 [species]
Onthophagus diversus Reiche, 1847 [species]
Onthophagus dives Harold, 1877 [species]
Onthophagus dlabolai Balthasar, 1964 [species]
Onthophagus dohertyi D'orbigny, 1902 [species]
Onthophagus doiinthanonensis Masumoto, 1989 [species]
Onthophagus doipuiensis Masumoto, 1989 [species]
Onthophagus doisuthepensis Masumoto, 1989 [species]
Onthophagus doitungensis Masumoto & Ochi & Hanboonsong, 2002 [species]
Onthophagus doriae Harold, 1877 [species]
Onthophagus dorsipilulus HOWDEN & GILL, 1987 [species]
Onthophagus dorsofasciatus Fairmaire, 1893 [species]
Onthophagus dorsosignatus D'Orbigny, 1898 [species]
Onthophagus dorsuosus D'orbigny, 1902 [species]
Onthophagus drescheri Paulian, 1939 [species]
Onthophagus dubernardi Boucomont, 1914 [species]
Onthophagus dubitabilis Howden & Génier, 2004 [species]
Onthophagus duboulayi Waterhouse, 1894 [species]
Onthophagus ducorpsi D'orbigny, 1913 [species]
Onthophagus dudleyi Cambefort, 1980 [species]
Onthophagus dummal Matthews, 1972 [species]
Onthophagus dunningi Harold, 1869 [species]
Onthophagus duporti Boucomont, 1914 [species]
Onthophagus duvivieri D'orbigny, 1904 [species]
Onthophagus dux Sharp, 1875 [species]
Onthophagus dynastoides Arrow, 1931 [species]
Onthophagus ebenicolor D'orbigny, 1902 [species]
Onthophagus ebenus Péringuey, 1888 [species]
Onthophagus eburneus D'orbigny, 1913 [species]
Onthophagus echinus Boucomont, 1914 [species]
Onthophagus ecopas Josso & Prévost, 2006 [species]
Onthophagus egenus Harold, 1877 [species]
Onthophagus egregius Arrow, 1907 [species]
Onthophagus egurianus Matsumura, 1938 [species]
Onthophagus elegans Klug, 1832 [species]
Onthophagus elgoni D'Orbigny, 1915 [species]
Onthophagus eliptaminus Balthasar, 1969 [species]
Onthophagus elongatus Frey, 1954 [species]
Onthophagus emarginatus Mulsant, 1842 [species]
Onthophagus embersoni Masumoto, Hanboonsong & Ochi, 2002 [species]
Onthophagus embrikianus Paulian, 1936 [species]
Onthophagus emeritus Péringuey, 1901 [species]
Onthophagus endota Matthews, 1972 [species]
Onthophagus endroedianus WALTER & CAMBEFORT, 1977 [species]
Onthophagus endroedii Frey, 1973 [species]
Onthophagus ensifer Boucomont, 1914 [species]
Onthophagus enuguensis Frey, 1975 [species]
Onthophagus ephippioderus Arrow, 1907 [species]
Onthophagus epilamprus Bates, 1888 [species]
Onthophagus erectinasus D'orbigny, 1902 [species]
Onthophagus erichsoni Hope, 1841 [species]
Onthophagus escalerai D'orbigny, 1902 [species]
Onthophagus eschscholtzi Boucomont, 1924 [species]
Onthophagus euchlorus D'Orbigny, 1902 [species]
Onthophagus eulaminicornis Pittino, 2006 [species]
Onthophagus eulophus Bates, 1887 [species]
Onthophagus euryceros Kabakov, 1998 [species]
Onthophagus euzeti WALTER & CAMBEFORT, 1977 [species]
Onthophagus evae Balthasar, 1944 [species]
Onthophagus evanidus Harold, 1869 [species]
Onthophagus exasperatus Gerstaecker, 1871 [species]
Onthophagus excisiceps D'orbigny, 1902 [species]
Onthophagus excisus Reiche & Saulcy, 1856 [species]
Onthophagus excubitor Ziani & Gudenzi, 2006 [species]
Onthophagus exiguus Raffray, 1877 [species]
Onthophagus exilis D'orbigny, 1913 [species]
Onthophagus expansicornis Bates, 1891 [species]
Onthophagus exquisitus Arrow, 1931 [species]
Onthophagus extensicollis D'orbigny, 1907 [species]
Onthophagus extensus Harold, 1878 [species]
Onthophagus fabricii Waterhouse, 1894 [species]
Onthophagus falcarius Balthasar, 1966 [species]
Onthophagus falcifer Harold, 1880 [species]
Onthophagus falculatus Boucomont, 1914 [species]
Onthophagus fallaciosus Raffray, 1877 [species]
Onthophagus fallax D'orbigny, 1913 [species]
Onthophagus falsivigilans Masumoto, 1995 [species]
Onthophagus falsus Gillet, 1925 [species]
Onthophagus falzonii Goidanich, 1926 [species]
Onthophagus fasciatus Boucomont, 1914 [species]
Onthophagus fasciculiger D'orbigny, 1902 [species]
Onthophagus fasciolatus Boucomont, 1924 [species]
Onthophagus fastosus D'Orbigny, 1913 [species]
Onthophagus favrei Boucomont, 1914 [species]
Onthophagus feai D'orbigny, 1905 [species]
Onthophagus felix Arrow, 1931 [species]
Onthophagus ferox Harold, 1867 [species]
Onthophagus ferrari Matthews, 1972 [species]
Onthophagus filicornis Harold, 1873 [species]
Onthophagus fimetarius Roth, 1851 [species]
Onthophagus finschi Harold, 1877 [species]
Onthophagus fissiceps Macleay, 1888 [species]
Onthophagus fissicornis Steven, 1809 [species]
Onthophagus fissinasus Fairmaire, 1895 [species]
Onthophagus fitiniensis Cambefort, 1984 [species]
Onthophagus flagrans Reitter, 1893 [species]
Onthophagus flavibasis D'orbigny, 1904 [species]
Onthophagus flaviclava D'orbigny, 1902 [species]
Onthophagus flavicornis Germar, 1824 [species]
Onthophagus flavimargo D'orbigny, 1902 [species]
Onthophagus flavipennis D'orbigny, 1905 [species]
Onthophagus flavoapicalis Lea, 1923 [species]
Onthophagus flavocinctus Frey, 1975 [species]
Onthophagus flavolimbatus Klug, 1855 [species]
Onthophagus flavomaculatus Gillet, 1930 [species]
Onthophagus flavorufus D'orbigny, 1904 [species]
Onthophagus fletcheri Blackburn, 1903 [species]
Onthophagus flexicollis D'Orbigny, 1904 [species]
Onthophagus flexicornis D'orbigny, 1902 [species]
Onthophagus flexifrons D'orbigny, 1908 [species]
Onthophagus fodiens Waterhouse, 1875 [species]
Onthophagus foedus Boucomont, 1914 [species]
Onthophagus foliaceus Lansberge, 1886 [species]
Onthophagus foliiceps Quedenfeldt, 1884 [species]
Onthophagus foraminosus D'orbigny, 1902 [species]
Onthophagus formaneki Reitter, 1897 [species]
Onthophagus formosanus Gillet, 1924 [species]
Onthophagus fortigibber Reitter, 1909 [species]
Onthophagus fossibasis D'orbigny, 1908 [species]
Onthophagus fossidorsis D'Orbigny, 1905 [species]
Onthophagus fossifrons D'orbigny, 1902 [species]
Onthophagus fossor Arrow, 1931 [species]
Onthophagus fossulatus D'orbigny, 1908 [species]
Onthophagus foulliouxi Cambefort, 1971 [species]
Onthophagus foveatus Frey, 1957 [species]
Onthophagus foveicollis Boucomont, 1930 [species]
Onthophagus fracticornis Preyssler, 1790 [species]
Onthophagus fradei Gomes Alves, 1956 [species]
Onthophagus frankenbergeri Balthasar, 1933 [species]
Onthophagus frankrozendaali Huijbregts & Krikken, 2012 [species]
Onthophagus frenchi Blackburn, 1903 [species]
Onthophagus freyi Moretto, 2004 [species]
Onthophagus fritschi D'orbigny, 1902 [species]
Onthophagus frontalis Raffray, 1877 [species]
Onthophagus frugivorus Arrow, 1931 [species]
Onthophagus fugitivus Péringuey, 1901 [species]
Onthophagus fujiii Ochi & Kon, 1995 [species]
Onthophagus fujiokai Ochi & Araya, 1996 [species]
Onthophagus fuliginosus Erichson, 1842 [species]
Onthophagus fulvocinctus D'orbigny, 1902 [species]
Onthophagus fulvus Sharp, 1875 [species]
Onthophagus fumatus D'Orbigny, 1913 [species]
Onthophagus funebris Boucomont, 1919 [species]
Onthophagus funestus Moretto & Génier, 2010 [species]
Onthophagus fungicola D'orbigny, 1908 [species]
Onthophagus furcaticeps Masters, 1886 [species]
Onthophagus furcatoides Lansberge, 1886 [species]
Onthophagus furcatus Fabricius, 1781 [species]
Onthophagus furciceps Marseul, 1869 [species]
Onthophagus furcicollis Arrow, 1931 [species]
Onthophagus furcifer Boheman, 1860 [species]
Onthophagus furcillifer Bates, 1891 [species]
Onthophagus furciramus D'Orbigny, 1902 [species]
Onthophagus furculifer D'orbigny, 1905 [species]
Onthophagus furculus Fabricius, 1798 [species]
Onthophagus fuscatus D'orbigny, 1908 [species]
Onthophagus fuscidorsis D ORBIGNY, 1902 [species]
Onthophagus fuscivestis D'orbigny, 1902 [species]
Onthophagus fuscopunctatus Fabricius, 1798 [species]
Onthophagus fuscostriatus Boucomont, 1914 [species]
Onthophagus gabonensis Walter, 1989 [species]
Onthophagus gaesatus Boucomont, 1925 [species]
Onthophagus gagates Hope, 1831 [species]
Onthophagus gagatinus Gillet, 1930 [species]
Onthophagus gagatoides Kabakov, 2006 [species]
Onthophagus gaillardi D'orbigny, 1911 [species]
Onthophagus gajo Krikken, 1977 [species]
Onthophagus galeatus Boucomont, 1919 [species]
Onthophagus ganalensis Gestro, 1895 [species]
Onthophagus gandju Matthews, 1972 [species]
Onthophagus gangeticus Gillet, 1925 [species]
Onthophagus gangloffi Josso & Prévost, 2001 [species]
Onthophagus gangulu Matthews, 1972 [species]
Onthophagus garambae Frey, 1961 [species]
Onthophagus gazellinus Bates, 1887 [species]
Onthophagus geelongensis Blackburn, 1891 [species]
Onthophagus gemellatus Raffray, 1877 [species]
Onthophagus geminatus D'orbigny, 1907 [species]
Onthophagus geminifrons D'orbigny, 1905 [species]
Onthophagus gemma Sharp, 1875 [species]
Onthophagus gemmatus Péringuey, 1901 [species]
Onthophagus genuinus Kohlmann & Solis, 2001 [species]
Onthophagus germanus Gillet, 1927 [species]
Onthophagus gerstaeckeri Harold, 1867 [species]
Onthophagus geryon Balthasar, 1969 [species]
Onthophagus gestroi Harold, 1877 [species]
Onthophagus ghanensis Balthasar, 1966 [species]
Onthophagus gibber D'orbigny, 1905 [species]
Onthophagus gibberosus D'orbigny, 1905 [species]
Onthophagus gibbicollis Lansberge, 1885 [species]
Onthophagus gibbidorsis D'orbigny, 1902 [species]
Onthophagus gibbifrons D'orbigny, 1902 [species]
Onthophagus gibbiramus D'Orbigny, 1902 [species]
Onthophagus gibbulus Pallas, 1781 [species]
Onthophagus gibbus D'orbigny, 1913 [species]
Onthophagus gibsoni Howden & Génier, 2004 [species]
Onthophagus gidju Matthews, 1972 [species]
Onthophagus gigantivigilans Masumoto, Hanboonsong & Ochi, 2002 [species]
Onthophagus gilli Delgado & Howden, 2000 [species]
Onthophagus ginyunensis Vsetecka, 1942 [species]
Onthophagus giraffa Hausmann, 1807 [species]
Onthophagus girardinae Walter, 1989 [species]
Onthophagus giuseppecarpanetoi Tagliaferri & Moretto, 2012 [species]
Onthophagus glabratus Hope, 1842 [species]
Onthophagus gladiator Arrow, 1907 [species]
Onthophagus glasunovi Koshantschikov, 1894 [species]
Onthophagus glaucinus Gillet, 1930 [species]
Onthophagus gnu Frey, 1955 [species]
Onthophagus godarra STOREY & WEIR, 1990 [species]
Onthophagus gonipa Krikken & Huijbregts, 2011 [species]
Onthophagus gonopygus D'orbigny, 1902 [species]
Onthophagus gorochovi Kabakov, 1998 [species]
Onthophagus gorodinskii Kabakov, 2008 [species]
Onthophagus gorokae Paulian, 1972 [species]
Onthophagus gosoli Masumoto, 1989 [species]
Onthophagus gothicus Gillet, 1930 [species]
Onthophagus gracilipes Boucomont, 1914 [species]
Onthophagus gradivus Balthasar, 1966 [species]
Onthophagus grandidorsis D'orbigny, 1913 [species]
Onthophagus grandifrons D'orbigny, 1907 [species]
Onthophagus grandivigilans Masumoto, 1995 [species]
Onthophagus graniceps D'orbigny, 1908 [species]
Onthophagus granosus D'orbigny, 1913 [species]
Onthophagus granulatus Boheman, 1858 [species]
Onthophagus granulifer Harold, 1886 [species]
Onthophagus granulifrons Frey, 1961 [species]
Onthophagus granulipennis Lansberge, 1886 [species]
Onthophagus granulum D'orbigny, 1904 [species]
Onthophagus granum Lansberge, 1885 [species]
Onthophagus graphicus Wallengren, 1881 [species]
Onthophagus grassei Walter, 1989 [species]
Onthophagus grataehelenae Kohlmann & Solis, 2001 [species]
Onthophagus gratus Arrow, 1931 [species]
Onthophagus gravearmatus Balthasar, 1944 [species]
Onthophagus gravis Walker, 1858 [species]
Onthophagus gravoti D'orbigny, 1908 [species]
Onthophagus griseoaeneus Lansberge, 1885 [species]
Onthophagus griseosetosus Arrow, 1931 [species]
Onthophagus grossepunctatus Reitter, 1905 [species]
Onthophagus guatemalensis Bates, 1887 [species]
Onthophagus gulmarri Matthews, 1972 [species]
Onthophagus gulo Arrow, 1931 [species]
Onthophagus gunsalami Ochi & Kon, 2005 [species]
Onthophagus gurburra Storey & Weir, 1990 [species]
Onthophagus guttatus Boheman, 1860 [species]
Onthophagus guttiger D'orbigny, 1905 [species]
Onthophagus haafi Frey, 1963 [species]
Onthophagus haagi Harold, 1867 [species]
Onthophagus haematopus Harold, 1875 [species]
Onthophagus hageni Lansberge, 1883 [species]
Onthophagus hajimei Masumoto, 1984 [species]
Onthophagus halffteri Zunino, 1981 [species]
Onthophagus hamaticeps Arrow, 1931 [species]
Onthophagus hamaticornis Gillet, 1930 [species]
Onthophagus haroldi Ballion, 1870 [species]
Onthophagus harpax Fabricius, 1801 [species]
Onthophagus hartiniae Ochi & Kon, 2009 [species]
Onthophagus hastifer Lansberge, 1885 [species]
Onthophagus hayashii Masumoto, 1991 [species]
Onthophagus hayashimasaoi Ochi & Kon, 1999 [species]
Onthophagus helciatus Harold, 1871 [species]
Onthophagus helmsi Blackburn, 1903 [species]
Onthophagus hemichalceus D'orbigny, 1915 [species]
Onthophagus hemichlorus D'orbigny, 1915 [species]
Onthophagus hemicyanus D'Orbigny, 1902 [species]
Onthophagus hemipygus Frey, 1964 [species]
Onthophagus herciniformis Moretto, 2010 [species]
Onthophagus hericius D'orbigny, 1902 [species]
Onthophagus hermonensis Baraud, 1982 [species]
Onthophagus herus Péringuey, 1901 [species]
Onthophagus heteroclitus D'orbigny, 1911 [species]
Onthophagus heterorrhinus Lansberge, 1885 [species]
Onthophagus heurni Gillet, 1930 [species]
Onthophagus heydeni Harold, 1875 [species]
Onthophagus heyrovskyi Vsetecka, 1943 [species]
Onthophagus hiabunicus Kabakov, 1998 [species]
Onthophagus hidakai Ochi & Kon, 1995 [species]
Onthophagus hielkemai Krikken & Huijbregts, 2011 [species]
Onthophagus hikidai Ochi & Kon, 2006 [species]
Onthophagus hilaridis Cambefort, 1984 [species]
Onthophagus hilaris D'orbigny, 1905 [species]
Onthophagus hildebrandti Harold, 1878 [species]
Onthophagus hindu Arrow, 1931 [species]
Onthophagus hingstoni Arrow, 1931 [species]
Onthophagus hippopotamus Harold, 1869 [species]
Onthophagus hirculus Mannerheim, 1829 [species]
Onthophagus hircus Billberg, 1815 [species]
Onthophagus hiroyukii Ochi, 2007 [species]
Onthophagus hirsutulus Lansberge, 1883 [species]
Onthophagus hirsutus D'orbigny, 1902 [species]
Onthophagus hirtellus Frey, 1957 [species]
Onthophagus hirticulus d'Orbigny, 1915 [species]
Onthophagus hirtipodex D'orbigny, 1904 [species]
Onthophagus hirtuosus Gillet, 1930 [species]
Onthophagus hirtus Illiger, 1803 [species]
Onthophagus hispanicus Baraud, 1963 [species]
Onthophagus hissariensis Kabakov, 2006 [species]
Onthophagus histeriformis Boucomont, 1914 [species]
Onthophagus histrio D'orbigny, 1902 [species]
Onthophagus hoberlandti Balthasar, 1966 [species]
Onthophagus holosericeus Harold, 1877 [species]
Onthophagus holzi Boucomont, 1914 [species]
Onthophagus hoogstraali Saylor, 1943 [species]
Onthophagus hoplocerus Lea, 1923 [species]
Onthophagus hoplothorax Gillet, 1930 [species]
Onthophagus horii Ochi, Kon & Tsubaki, 2009 [species]
Onthophagus horni Balthasar, 1935 [species]
Onthophagus horrens D'orbigny, 1908 [species]
Onthophagus horribilis Balthasar, 1969 [species]
Onthophagus horridus D'orbigny, 1908 [species]
Onthophagus howdenorum ZUNINO & HALFFTER, 1988 [species]
Onthophagus hsui Masumoto, Chen & Ochi, 2004 [species]
Onthophagus hulstaerti Boucomont, 1932 [species]
Onthophagus humpatensis Lansberge, 1886 [species]
Onthophagus hyaena Fabricius, 1801 [species]
Onthophagus hyalcyon Palestrini & Giacone, 1989 [species]
Onthophagus hystrix Boucomont, 1914 [species]
Onthophagus ibex Fabricius, 1792 [species]
Onthophagus ieti Cambefort, 1984 [species]
Onthophagus ifugaoensis Ochi & Kon, 2006 [species]
Onthophagus igneus Vigors, 1825 [species]
Onthophagus ignitus D'Orbigny, 1905 [species]
Onthophagus igualensis Bates, 1887 [species]
Onthophagus illotus Péringuey, 1901 [species]
Onthophagus illyricus Scopoli, 1763 [species]
Onthophagus imbellis D'orbigny, 1905 [species]
Onthophagus imberbis D'orbigny, 1902 [species]
Onthophagus imbutus Sharp, 1875 [species]
Onthophagus immundus Boheman, 1858 [species]
Onthophagus impar Kabakov, 2006 [species]
Onthophagus imperator Castelnau, 1840 [species]
Onthophagus importunus Péringuey, 1901 [species]
Onthophagus impressicollis Boheman, 1860 [species]
Onthophagus impunctatus D'orbigny, 1904 [species]
Onthophagus impurus Harold, 1868 [species]
Onthophagus inaequalis D'orbigny, 1902 [species]
Onthophagus incantatus Balthasar, 1967 [species]
Onthophagus incanus Macleay, 1888 [species]
Onthophagus incertus D'Orbigny, 1897 [species]
Onthophagus incisus Harold, 1877 [species]
Onthophagus includens D'orbigny, 1905 [species]
Onthophagus inclusus D'orbigny, 1905 [species]
Onthophagus incollaris Kabakov, 2008 [species]
Onthophagus incornutus Macleay, 1871 [species]
Onthophagus incostatus D'Orbigny, 1902 [species]
Onthophagus indigus Péringuey, 1901 [species]
Onthophagus indosinicus Boucomont, 1921 [species]
Onthophagus indutus D'orbigny, 1902 [species]
Onthophagus inediapterus Kohlmann & Solis, 2001 [species]
Onthophagus inelegans Balthasar, 1935 [species]
Onthophagus ineptus Harold, 1871 [species]
Onthophagus inermiceps D'orbigny, 1905 [species]
Onthophagus inermicollis D'orbigny, 1908 [species]
Onthophagus inermivertex Boucomont, 1921 [species]
Onthophagus infaustus Cambefort, 1984 [species]
Onthophagus inflaticollis Bates, 1887 [species]
Onthophagus inflatus WALTER & CAMBEFORT, 1977 [species]
Onthophagus infucatus Harold, 1877 [species]
Onthophagus infuscatus Klug, 1845 [species]
Onthophagus insignicollis Frey, 1954 [species]
Onthophagus insignis Péringuey, 1896 [species]
Onthophagus insularis Boheman, 1858 [species]
Onthophagus insulindicus Boucomont, 1914 [species]
Onthophagus insulsus Péringuey, 1901 [species]
Onthophagus intermixtus D'orbigny, 1902 [species]
Onthophagus interruptus Raffray, 1877 [species]
Onthophagus interstitialis Fahraeus, 1857 [species]
Onthophagus intonsus D'orbigny, 1913 [species]
Onthophagus intricatus Moretto, 2010 [species]
Onthophagus investigator Lansberge, 1885 [species]
Onthophagus investis D'orbigny, 1904 [species]
Onthophagus iodiellus Bates, 1887 [species]
Onthophagus irianus Balthasar, 1969 [species]
Onthophagus iris Sharp, 1875 [species]
Onthophagus isanus Masumoto, Ochi & Hanboonsong, 2008 [species]
Onthophagus ishiii OCHI & KON, 1995 [species]
Onthophagus isikdagensis Pittino, 2004 [species]
Onthophagus itiokai Ochi, Kon & Kishimoto-Yamada, 2010 [species]
Onthophagus itoi Nomura, 1976 [species]
Onthophagus iulicola Cambefort, 1984 [species]
Onthophagus iumienus Masumoto, Ochi & Hanboonsong, 2007 [species]
Onthophagus iyengari Arrow, 1931 [species]
Onthophagus jacksoni D'orbigny, 1913 [species]
Onthophagus jacobeus Boucomont, 1924 [species]
Onthophagus jalamari Matthews, 1972 [species]
Onthophagus jalapensis Balthasar, 1939 [species]
Onthophagus jangga Matthews, 1972 [species]
Onthophagus janssensi Frey, 1958 [species]
Onthophagus janthinus Harold, 1875 [species]
Onthophagus janushevi Kabakov, 1983 [species]
Onthophagus japonicus Harold, 1875 [species]
Onthophagus javacupreus Krikken & Huijbregts, 2009 [species]
Onthophagus javaecola Balthasar, 1959 [species]
Onthophagus javanensis Balthasar, 1963 [species]
Onthophagus javanitidus Ochi, Kon & Hartini, 2010 [species]
Onthophagus javanoides Huijbregts & Krikken, 2011 [species]
Onthophagus jeanneli D'orbigny, 1913 [species]
Onthophagus jeannelianus Paulian, 1945 [species]
Onthophagus jingping Masumoto, Ochi & Hanboonsong, 2007 [species]
Onthophagus joannae Goljan, 1953 [species]
Onthophagus johkii Ochi & Kon, 1994 [species]
Onthophagus joliveti Paulian, 1972 [species]
Onthophagus jubatus Harold, 1869 [species]
Onthophagus jugicola D'orbigny, 1902 [species]
Onthophagus juncticornis D'orbigny, 1908 [species]
Onthophagus justei Walter, 1989 [species]
Onthophagus juvencus Klug, 1835 [species]
Onthophagus kabakovi Martin-Piera, 1985 [species]
Onthophagus kabulicus Balthasar, 1971 [species]
Onthophagus kachinicus Kabakov, 2006 [species]
Onthophagus kachowskii Olsoufief, 1900 [species]
Onthophagus kaengkrachangus Masumoto, Ochi & Hanboonsong, 2008 [species]
Onthophagus kakadu STOREY & WEIR, 1990 [species]
Onthophagus kanarensis Arrow, 1931 [species]
Onthophagus kangeanus Paulian, 1936 [species]
Onthophagus kanyaayonus Masumoto, 1992 [species]
Onthophagus kaosoidowensis Masumoto, Ochi & Hanboonsong, 2008 [species]
Onthophagus kapitensis Frey, 1971 [species]
Onthophagus kapuri Endrödi, 1974 [species]
Onthophagus karenensis Masumoto, Ochi & Hanboonsong, 2008 [species]
Onthophagus kashizakii Kon & Ochi, 2005 [species]
Onthophagus kashmirensis Balthasar, 1966 [species]
Onthophagus kassaicus D'orbigny, 1908 [species]
Onthophagus katangensis Frey, 1961 [species]
Onthophagus katoi Ochi & Araya, 1996 [species]
Onthophagus katualensis Quedenfeldt, 1888 [species]
Onthophagus kavirondus D'orbigny, 1915 [species]
Onthophagus kawaharai Ochi & Kon, 2007 [species]
Onthophagus kchatriya Boucomont, 1914 [species]
Onthophagus keiseri Frey, 1956 [species]
Onthophagus kentingensis Nomura, 1973 [species]
Onthophagus keralensis Frey, 1975 [species]
Onthophagus keralicus Biswas & Chatterjee, 1986 [species]
Onthophagus khonkaenus Masumoto, Ochi & Hanboonsong, 2008 [species]
Onthophagus khonmiinitnoi Masumoto, 1990 [species]
Onthophagus kiambram Storey, 1977 [species]
Onthophagus kikutai Ochi & Kon, 2005 [species]
Onthophagus kilimanus Kolbe, 1891 [species]
Onthophagus kimioi Ochi, Kon & Kawahara, 2011 [species]
Onthophagus kindermanni Harold, 1877 [species]
Onthophagus kindianus Frey, 1953 [species]
Onthophagus kingstoni Cambefort, 1980 [species]
Onthophagus kinhthaicus Kabakov, 1998 [species]
Onthophagus kirki Frey, 1975 [species]
Onthophagus kirokanus Frey, 1975 [species]
Onthophagus kiuchianus Masumoto, Yang & Ochi, 2004 [species]
Onthophagus kiuchii Masumoto, 1995 [species]
Onthophagus kiyoshii Ochi & Kon, 2007 [species]
Onthophagus klapperichi Balthasar, 1952 [species]
Onthophagus kleinei Balthasar, 1935 [species]
Onthophagus knapperti Krikken, 1981 [species]
Onthophagus kochi Frey, 1957 [species]
Onthophagus koebelei Blackburn, 1903 [species]
Onthophagus koechlei Masumoto, Ochi & Kon, 2011 [species]
Onthophagus koichii Ochi & Araya, 1992 [species]
Onthophagus kokereka Matthews, 1972 [species]
Onthophagus kolaka Huijbregts & Krikken, 2009 [species]
Onthophagus kolbei Marcus, 1921 [species]
Onthophagus kolenatii Reitter, 1893 [species]
Onthophagus koma Matsumura, 1937 [species]
Onthophagus komareki Balthasar, 1935 [species]
Onthophagus kondaoensis Kabakov, 1994 [species]
Onthophagus koni Ochi, 2007 [species]
Onthophagus konoi Matsumura, 1938 [species]
Onthophagus konsarnensis Masumoto, Ochi & Hanboonsong, 2008 [species]
Onthophagus kontumicus Kabakov, 1983 [species]
Onthophagus kora Storey, 1977 [species]
Onthophagus koryoensis Kim, 1985 [species]
Onthophagus koshunensis Balthasar, 1941 [species]
Onthophagus kouassii Cambefort, 1984 [species]
Onthophagus kozlovi Kabakov, 1990 [species]
Onthophagus kraatzeanus Lansberge, 1883 [species]
Onthophagus krakadaakhomus Masumoto, 1992 [species]
Onthophagus kryzhanovskii Kabakov, 1982 [species]
Onthophagus kuatunensis Balthasar, 1942 [species]
Onthophagus kukunorensis Kabakov, 1990 [species]
Onthophagus kulti Balthasar, 1952 [species]
Onthophagus kuluensis Bates, 1891 [species]
Onthophagus kulzeri Frey, 1958 [species]
Onthophagus kumaonensis Arrow, 1931 [species]
Onthophagus kumbaingeri Matthews, 1972 [species]
Onthophagus kwaleensis Moretto & Nicolas, 2002 [species]
Onthophagus kyleensis Frey, 1975 [species]
Onthophagus labdacus Balthasar, 1969 [species]
Onthophagus laborans Arrow, 1931 [species]
Onthophagus laceratus Gerstaecker, 1871 [species]
Onthophagus lacustris Harold, 1877 [species]
Onthophagus laetus D'orbigny, 1913 [species]
Onthophagus laevatus D'orbigny, 1902 [species]
Onthophagus laeviceps D'orbigny, 1902 [species]
Onthophagus laevicollis Lansberge, 1883 [species]
Onthophagus laevidorsis D'orbigny, 1907 [species]
Onthophagus laevigatus Fabricius, 1798 [species]
Onthophagus laevissimus D'orbigny, 1905 [species]
Onthophagus lagnyi Paulian, 1937 [species]
Onthophagus lahorensis Kabakov, 2008 [species]
Onthophagus lakyim Masumoto, 1990 [species]
Onthophagus lallieri d'Orbigny, 1902 [species]
Onthophagus lallierianus Janssens, 1953 [species]
Onthophagus lamellicornis Frey, 1953 [species]
Onthophagus lamelliger Gerstaecker, 1871 [species]
Onthophagus lamgalio Matthews, 1972 [species]
Onthophagus laminatus Macleay, 1864 [species]
Onthophagus laminicornis Lansberge, 1886 [species]
Onthophagus laminidorsis D'orbigny, 1902 [species]
Onthophagus laminosus D'orbigny, 1905 [species]
Onthophagus lamnifer D'orbigny, 1902 [species]
Onthophagus lamotellus Cambefort, 1980 [species]
Onthophagus lamottei Frey, 1962 [species]
Onthophagus lamtoensis Cambefort, 1984 [species]
Onthophagus lamtoi Cambefort, 1980 [species]
Onthophagus lamyi Paulian, 1945 [species]
Onthophagus lanista Castelnau, 1840 [species]
Onthophagus lannamiibun Masumoto, Ochi & Hanboonsong, 2002 [species]
Onthophagus laotianus Boucomont, 1919 [species]
Onthophagus lapillus Arrow, 1931 [species]
Onthophagus laratinus Arrow, 1916 [species]
Onthophagus lassulus Balthasar, 1964 [species]
Onthophagus latenasutus Arrow, 1941 [species]
Onthophagus latepunctatus D'orbigny, 1913 [species]
Onthophagus latestriatus D'orbigny, 1908 [species]
Onthophagus latevittatus D'orbigny, 1905 [species]
Onthophagus laticollis Klug, 1835 [species]
Onthophagus laticornis Gebler, 1823 [species]
Onthophagus latigena D'Orbigny, 1897 [species]
Onthophagus latigibber D ORBIGNY, 1902 [species]
Onthophagus latipennis D'orbigny, 1897 [species]
Onthophagus latissimus D'orbigny, 1902 [species]
Onthophagus latro Harold, 1877 [species]
Onthophagus leai Blackburn, 1895 [species]
Onthophagus leanus Goidanich, 1926 [species]
Onthophagus lebasi Boucomont, 1932 [species]
Onthophagus lefebvrei D'orbigny, 1913 [species]
Onthophagus lefiniensis Balthasar, 1967 [species]
Onthophagus legendrei WALTER & CAMBEFORT, 1977 [species]
Onthophagus lemagneni D'orbigny, 1902 [species]
Onthophagus lemekensis D'orbigny, 1915 [species]
Onthophagus lemniscatus Gillet, 1924 [species]
Onthophagus lemur Fabricius, 1781 [species]
Onthophagus lemuroides D'orbigny, 1898 [species]
Onthophagus lenis Kabakov, 1998 [species]
Onthophagus lenzi Harold, 1875 [species]
Onthophagus leomontanus Boucomont, 1914 [species]
Onthophagus leroyi D'orbigny, 1902 [species]
Onthophagus leucopygus Harold, 1867 [species]
Onthophagus leucostigma Steven, 1811 [species]
Onthophagus leusermontis Huijbregts & Krikken, 2011 [species]
Onthophagus leytensis Ochi & Araya, 1992 [species]
Onthophagus liberianus Lansberge, 1883 [species]
Onthophagus liewi Ochi & Kon, 2005 [species]
Onthophagus lilliputanus Lansberge, 1883 [species]
Onthophagus limbatus Herbst, 1789 [species]
Onthophagus limonensis Kohlmann & Solis, 2001 [species]
Onthophagus lindaae Masumoto, 1989 [species]
Onthophagus lindu Huijbregts & Krikken, 2009 [species]
Onthophagus liodermus D'orbigny, 1913 [species]
Onthophagus lioides D'orbigny, 1902 [species]
Onthophagus liopterus Harold, 1880 [species]
Onthophagus liothorax Koshantschikov, 1894 [species]
Onthophagus lituratus Roth, 1851 [species]
Onthophagus liwagensis Ochi & Kon, 2006 [species]
Onthophagus lobi Cambefort, 1984 [species]
Onthophagus lojanus Balthasar, 1939 [species]
Onthophagus lomii Müller, 1942 [species]
Onthophagus longefossus D'Orbigny, 1905 [species]
Onthophagus longegranus Frey, 1957 [species]
Onthophagus longiceps D'orbigny, 1904 [species]
Onthophagus longimanus Bates, 1887 [species]
Onthophagus longipes Paulian, 1937 [species]
Onthophagus longipilis D'orbigny, 1905 [species]
Onthophagus lore Huijbregts & Krikken, 2009 [species]
Onthophagus lorianus Gillet, 1930 [species]
Onthophagus loricatus Klug, 1855 [species]
Onthophagus loroi Balthasar, 1941 [species]
Onthophagus loudetiae Cambefort, 1984 [species]
Onthophagus loveni Gillet, 1928 [species]
Onthophagus loxodontae Cambefort, 1984 [species]
Onthophagus loxodontaphilus Moretto, 2009 [species]
Onthophagus lucasi D'Orbigny, 1902 [species]
Onthophagus lucidus Illiger, 1800 [species]
Onthophagus luctuosus Boucomont, 1914 [species]
Onthophagus ludicrus Balthasar, 1969 [species]
Onthophagus ludio Boucomont, 1914 [species]
Onthophagus lugens Fahraeus, 1857 [species]
Onthophagus lugubris Fahraeus, 1857 [species]
Onthophagus luismargaritorum Delgado, 1995 [species]
Onthophagus lujendae Bates, 1888 [species]
Onthophagus lumareti WALTER & CAMBEFORT, 1977 [species]
Onthophagus lunatus Harold, 1868 [species]
Onthophagus lunulifer Boucomont, 1914 [species]
Onthophagus luridipennis Boheman, 1858 [species]
Onthophagus lutaticollis D'orbigny, 1907 [species]
Onthophagus luteosignatus Lansberge, 1883 [species]
Onthophagus lutosopictus Fairmaire, 1897 [species]
Onthophagus macedonicus Miksic, 1959 [species]
Onthophagus macleayi Blackburn, 1903 [species]
Onthophagus macrocephalus Kirby, 1818 [species]
Onthophagus macroliberianus Moretto, 2010 [species]
Onthophagus macrothorax D'orbigny, 1902 [species]
Onthophagus maculatus Fabricius, 1801 [species]
Onthophagus maculosipennis Gillet, 1930 [species]
Onthophagus maculosus D'orbigny, 1908 [species]
Onthophagus madoqua Arrow, 1931 [species]
Onthophagus maephaluangus Masumoto, Ochi & Hanboonsong, 2007 [species]
Onthophagus maesaensis Masumoto, Ochi & Hanboonsong, 2002 [species]
Onthophagus magnini Paulian, 1933 [species]
Onthophagus magnioculus Ochi & Kon, 2006 [species]
Onthophagus magnipygus Boucomont, 1914 [species]
Onthophagus mairuu Masumoto, 1989 [species]
Onthophagus maki Illiger, 1803 [species]
Onthophagus makokou Walter, 1989 [species]
Onthophagus malabarensis Boucomont, 1919 [species]
Onthophagus malangensis Boucomont, 1914 [species]
Onthophagus malasiacus Gillet, 1927 [species]
Onthophagus maleengnaafon Masumoto, 1990 [species]
Onthophagus maleengnoi Masumoto, 1990 [species]
Onthophagus malevolus Balthasar, 1964 [species]
Onthophagus malthinus Gillet, 1930 [species]
Onthophagus mamillatus Lea, 1923 [species]
Onthophagus manguliensis Boucomont, 1914 [species]
Onthophagus manipurensis Arrow, 1907 [species]
Onthophagus maniti Masumoto, 1989 [species]
Onthophagus mankonoensis Balthasar, 1966 [species]
Onthophagus manya Matthews, 1972 [species]
Onthophagus marahouensis Cambefort, 1984 [species]
Onthophagus margaretensis Blackburn, 1903 [species]
Onthophagus margaritifer D'orbigny, 1898 [species]
Onthophagus marginatus Castelnau, 1840 [species]
Onthophagus marginicollis Harold, 1880 [species]
Onthophagus marginidens D'Orbigny, 1904 [species]
Onthophagus marginifer Frey, 1953 [species]
Onthophagus maringouensis Palestrini, 1992 [species]
Onthophagus mariozuninoi Delgado, Navarrete & Blackaller-Bages, 1993 [species]
Onthophagus marmotae Kabakov, 1990 [species]
Onthophagus marshalli D'orbigny, 1908 [species]
Onthophagus martelli Frey, 1972 [species]
Onthophagus martialis Boucomont, 1914 [species]
Onthophagus maryatiae Ochi & Kon, 2005 [species]
Onthophagus masahiroi Masumoto, Ochi & Hanboonsong, 2008 [species]
Onthophagus masaicus D'orbigny, 1905 [species]
Onthophagus masaoi Ochi, 1992 [species]
Onthophagus massai Baraud, 1975 [species]
Onthophagus masumotoi Ochi, 1985 [species]
Onthophagus matae Cambefort, 1984 [species]
Onthophagus matanyo Huijbregts & Krikken, 2009 [species]
Onthophagus matsudai Ochi & Kon, 2004 [species]
Onthophagus matsuii Ochi & Kon, 2006 [species]
Onthophagus mauritii Boucomont, 1919 [species]
Onthophagus maya Zunino, 1981 [species]
Onthophagus mayeri Harold, 1876 [species]
Onthophagus mediofuscatus D'Orbigny, 1908 [species]
Onthophagus medius Kugelann, 1792 [species]
Onthophagus medvedevi Kabakov, 1982 [species]
Onthophagus megapacificus Ochi & Kon, 2006 [species]
Onthophagus megathorax Gillet, 1921 [species]
Onthophagus mekara Huijbregts & Krikken, 2009 [species]
Onthophagus melanocephalus Klug, 1845 [species]
Onthophagus melitaeus Fabricius, 1798 [species]
Onthophagus mendeli Ochi, Kon & Barclay, 2009 [species]
Onthophagus mendicus Gillet, 1924 [species]
Onthophagus menieri Ochi, 2003 [species]
Onthophagus menkaoensis WALTER & CAMBEFORT, 1977 [species]
Onthophagus mentaveiensis Boucomont, 1914 [species]
Onthophagus merdarius Chevrolat, 1865 [species]
Onthophagus merdrignaci WALTER & CAMBEFORT, 1977 [species]
Onthophagus meruanus D'orbigny, 1908 [species]
Onthophagus merus Péringuey, 1901 [species]
Onthophagus metalliceps Arrow, 1931 [species]
Onthophagus metalliger D'orbigny, 1913 [species]
Onthophagus mexicanus Bates, 1887 [species]
Onthophagus micropterus ZUNINO & HALFFTER, 1981 [species]
Onthophagus micros D'orbigny, 1902 [species]
Onthophagus mije STOREY & WEIR, 1990 [species]
Onthophagus miles D'orbigny, 1905 [species]
Onthophagus militaris Boucomont, 1914 [species]
Onthophagus millamilla Matthews, 1972 [species]
Onthophagus millingeni D'orbigny, 1898 [species]
Onthophagus mimikanus Balthasar, 1969 [species]
Onthophagus mimus D'orbigny, 1913 [species]
Onthophagus minans D'orbigny, 1913 [species]
Onthophagus minax D'orbigny, 1904 [species]
Onthophagus mindanaensis Boucomont, 1914 [species]
Onthophagus mindanaoensis Frey, 1971 [species]
Onthophagus minettii Moretto & Nicolas, 2002 [species]
Onthophagus minotaurus Arrow, 1941 [species]
Onthophagus minutissimus D'orbigny, 1908 [species]
Onthophagus minutulus Harold, 1875 [species]
Onthophagus minutus Hausmann, 1807 [species]
Onthophagus mirabilis Bates, 1887 [species]
Onthophagus mirandus Arrow, 1931 [species]
Onthophagus miricollis Frey, 1962 [species]
Onthophagus miricornis d'Orbigny, 1902 [species]
Onthophagus mirifrons D'orbigny, 1905 [species]
Onthophagus miscellaneus D'orbigny, 1908 [species]
Onthophagus miscellus D'orbigny, 1905 [species]
Onthophagus misellus D'orbigny, 1907 [species]
Onthophagus missor Balthasar, 1938 [species]
Onthophagus mixticeps D'orbigny, 1905 [species]
Onthophagus mixtidorsis D'orbigny, 1905 [species]
Onthophagus mixtifrons D'Orbigny, 1910 [species]
Onthophagus miyakei Ochi & Araya, 1992 [species]
Onthophagus mjobergi Gillet, 1925 [species]
Onthophagus mniszechi Harold, 1869 [species]
Onthophagus moajat Huijbregts & Krikken, 2009 [species]
Onthophagus mocquerysi D'orbigny, 1902 [species]
Onthophagus modestus Harold, 1862 [species]
Onthophagus mogo Krikken & Huijbregts, 2008 [species]
Onthophagus monardi Boucomont, 1936 [species]
Onthophagus monardiellus Frey, 1963 [species]
Onthophagus monforti Josso & Prévost, 2006 [species]
Onthophagus mongana STOREY & WEIR, 1990 [species]
Onthophagus mongkhoni Masumoto, Hanboonsong & Ochi, 2002 [species]
Onthophagus monochromus D'Orbigny, 1913 [species]
Onthophagus monodon Fahraeus, 1857 [species]
Onthophagus monteithi Matthews, 1972 [species]
Onthophagus monticupreus Krikken & Huijbregts, 2009 [species]
Onthophagus montishannoniae Krikken & Huijbrechts, 2008 [species]
Onthophagus montivagus D'orbigny, 1902 [species]
Onthophagus montreuili Moretto & Génier, 2010 [species]
Onthophagus mopsus Fabricius, 1792 [species]
Onthophagus mordindangus Masumoto, Ochi & Hanboonsong, 2008 [species]
Onthophagus moreleti Baraud, 1980 [species]
Onthophagus moroni ZUNINO & HALFFTER, 1988 [species]
Onthophagus morosus Gerstaecker, 1871 [species]
Onthophagus mouhoti Harold, 1875 [species]
Onthophagus mpassa Walter, 1982 [species]
Onthophagus mucronatus Thomson, 1858 [species]
Onthophagus mucronifer D'orbigny, 1905 [species]
Onthophagus muelleri Lansberge, 1883 [species]
Onthophagus mulgravei Paulian, 1937 [species]
Onthophagus multicornis D'Orbigny, 1908 [species]
Onthophagus multipunctatus Boucomont, 1914 [species]
Onthophagus mundill Matthews, 1972 [species]
Onthophagus murchisoni Blackburn, 1892 [species]
Onthophagus mushensis Matsumura, 1938 [species]
Onthophagus mutatus Harold, 1859 [species]
Onthophagus muticifrons Endrödi, 1973 [species]
Onthophagus muticus Macleay, 1864 [species]
Onthophagus naaroon Masumoto, 1990 [species]
Onthophagus nabeleki Balthasar, 1939 [species]
Onthophagus naevius D'orbigny, 1913 [species]
Onthophagus naevuliger D'orbigny, 1908 [species]
Onthophagus nagasawai Matsumura, 1938 [species]
Onthophagus nagpurensis Arrow, 1931 [species]
Onthophagus nammuldi Matthews, 1972 [species]
Onthophagus namnaoensis Masumoto & Ochi & Hanboonsong, 2002 [species]
Onthophagus nampatensis Tarasov & Kabakov, 2010 [species]
Onthophagus nanus Harold, 1878 [species]
Onthophagus napolovi Kabakov, 1998 [species]
Onthophagus nasalis Arrow, 1931 [species]
Onthophagus nasicornis Harold, 1869 [species]
Onthophagus nasicus Gillet, 1930 [species]
Onthophagus nasidens D'Orbigny, 1905 [species]
Onthophagus naso Fahraeus, 1857 [species]
Onthophagus nasonis Kabakov, 1998 [species]
Onthophagus nasutus Guérin-Méneville, 1855 [species]
Onthophagus natronis Cambefort, 1984 [species]
Onthophagus navarretorum DELGADO & CAPISTAN, 1996 [species]
Onthophagus naviculifer Boucomont, 1919 [species]
Onthophagus neboissi Frey, 1970 [species]
Onthophagus nebulosus Reiche, 1864 [species]
Onthophagus necessarius Reitter, 1893 [species]
Onthophagus necrophagus Arrow, 1931 [species]
Onthophagus neervoorti Boucomont, 1914 [species]
Onthophagus nefarius Balthasar, 1941 [species]
Onthophagus negligens Walker, 1858 [species]
Onthophagus negrosensis Ochi & Araya, 1992 [species]
Onthophagus negus Raffray, 1882 [species]
Onthophagus nemorivagus Kohlmann & Solis, 2001 [species]
Onthophagus neocolobus Scheuern, 1996 [species]
Onthophagus neofurcatus Goidanich, 1926 [species]
Onthophagus neomirabilis Howden, 1973 [species]
Onthophagus neostenocerus Goidanich, 1926 [species]
Onthophagus ngottoensis Moretto & Nicolas, 2002 [species]
Onthophagus niasensis Boucomont, 1914 [species]
Onthophagus nicobaricus Biswas, Chatterjee & Sengupta, 1999 [species]
Onthophagus nicolasi Moretto, 2004 [species]
Onthophagus nigellus Illiger, 1803 [species]
Onthophagus nigerianus D'orbigny, 1913 [species]
Onthophagus nigriceps Raffray, 1877 [species]
Onthophagus nigricornis Fairmaire, 1887 [species]
Onthophagus nigrinus Paulian, 1937 [species]
Onthophagus nigriobscurior Ochi, Kon & Tsubaki, 2009 [species]
Onthophagus nigripennis D'orbigny, 1908 [species]
Onthophagus nigriventris D'orbigny, 1905 [species]
Onthophagus nigrivestis D'Orbigny, 1902 [species]
Onthophagus nigropubens D'orbigny, 1908 [species]
Onthophagus nigroviolaceus d'Orbigny, 1902 [species]
Onthophagus nikolajevi Kabakov, 2006 [species]
Onthophagus nilgirensis Gillet, 1922 [species]
Onthophagus nilicola D'orbigny, 1902 [species]
Onthophagus niloticus Harold, 1879 [species]
Onthophagus nimbatus D'orbigny, 1905 [species]
Onthophagus niokolokoba Moretto, 2007 [species]
Onthophagus nitefactus Harold, 1877 [species]
Onthophagus nitidifrons D'orbigny, 1905 [species]
Onthophagus nitidior Bates, 1887 [species]
Onthophagus nitidulus Klug, 1845 [species]
Onthophagus nitidus Waterhouse, 1875 [species]
Onthophagus nodieri D'orbigny, 1908 [species]
Onthophagus nodulifer Harold, 1867 [species]
Onthophagus nongkaiensis Masumoto, Ochi & Hanboonsong, 2002 [species]
Onthophagus nonstriatus D'orbigny, 1905 [species]
Onthophagus notatus D'orbigny, 1902 [species]
Onthophagus notiodes Solis & Kohlmann, 2003 [species]
Onthophagus novaeirlandiae Balthasar, 1969 [species]
Onthophagus nuba D'Orbigny, 1902 [species]
Onthophagus nubilus Kohlmann & Solis, 2001 [species]
Onthophagus nudatus D'orbigny, 1913 [species]
Onthophagus nudifrons Balthasar, 1939 [species]
Onthophagus nudus D'orbigny, 1908 [species]
Onthophagus numidicus d'Orbigny, 1908 [species]
Onthophagus nurestanicus Kabakov, 1982 [species]
Onthophagus nurubuan Matthews, 1972 [species]
Onthophagus nusatenggaricus Krikken & Huijbreghts, 2008 [species]
Onthophagus nyctopus Bates, 1887 [species]
Onthophagus nymani Gillet, 1930 [species]
Onthophagus obenbergeri Balthasar, 1965 [species]
Onthophagus oberthuri D'orbigny, 1898 [species]
Onthophagus obliquus Olivier, 1789 [species]
Onthophagus obliviosus Balthasar, 1935 [species]
Onthophagus oblongus Zhang, 1997 [species]
Onthophagus obscurior Boucomont, 1914 [species]
Onthophagus obtusicornis Fahraeus, 1857 [species]
Onthophagus obtutus Péringuey, 1901 [species]
Onthophagus occipitalis Lansberge, 1885 [species]
Onthophagus ocellatopunctatus Waterhouse, 1875 [species]
Onthophagus ocellidorsis D'orbigny, 1915 [species]
Onthophagus ocellifer D'orbigny, 1902 [species]
Onthophagus ocelliger Harold, 1877 [species]
Onthophagus ochii Masumoto, 1988 [species]
Onthophagus ochreatus D'Orbigny, 1897 [species]
Onthophagus ochromerus Harold, 1877 [species]
Onthophagus ochropygus D'orbigny, 1902 [species]
Onthophagus octogonus Frey, 1962 [species]
Onthophagus octonaevus Kabakov, 1994 [species]
Onthophagus oculatus Arrow, 1931 [species]
Onthophagus ohbayashii Nomura, 1939 [species]
Onthophagus ohkuboi Ochi & Kon, 2006 [species]
Onthophagus ohmomoi Miyake, 2000 [species]
Onthophagus okahandjanus Balthasar, 1974 [species]
Onthophagus olidus Balthasar, 1964 [species]
Onthophagus olsoufieffi Boucomont, 1924 [species]
Onthophagus omostigma D'orbigny, 1902 [species]
Onthophagus onorei ZUNINO & HALFFTER, 1997 [species]
Onthophagus onthochromus Arrow, 1913 [species]
Onthophagus ontosatu Huijbregts & Krikken, 2012 [species]
Onthophagus opacicollis Reitter, 1893 [species]
Onthophagus opacifalculatus Ochi, Kon & Tsubaki, 2009 [species]
Onthophagus ophtalmicus Frey, 1958 [species]
Onthophagus opulentus D'Orbigny, 1902 [species]
Onthophagus orbicularis Lansberge, 1885 [species]
Onthophagus orbus Boucomont, 1924 [species]
Onthophagus orientalis Harold, 1868 [species]
Onthophagus orissanus Arrow, 1931 [species]
Onthophagus ornaticollis Gillet, 1930 [species]
Onthophagus ornatulus D'orbigny, 1908 [species]
Onthophagus orphnoides Bates, 1887 [species]
Onthophagus orthocerus Thomson, 1858 [species]
Onthophagus osculatii Guérin-Méneville, 1855 [species]
Onthophagus osellai Pittino, 1982 [species]
Onthophagus oshimanus Nakane, 1960 [species]
Onthophagus otai Ochi & Kon, 2006 [species]
Onthophagus otjivarongus Balthasar, 1967 [species]
Onthophagus ouratita Matthews, 1972 [species]
Onthophagus ovatus Linnaeus, 1767 [species]
Onthophagus overlaeti Janssens, 1953 [species]
Onthophagus ovigranosus Frey, 1971 [species]
Onthophagus ovulum Gerstaecker, 1871 [species]
Onthophagus pacificus Lansberge, 1885 [species]
Onthophagus pactolus Fabricius, 1787 [species]
Onthophagus padrinoi Delgado, 1999 [species]
Onthophagus palatus Boucomont, 1914 [species]
Onthophagus palavanus Balthasar, 1959 [species]
Onthophagus palawanicus Ochi & Kon, 2004 [species]
Onthophagus palestriniae Moretto & Nicolas, 2004 [species]
Onthophagus paliceps Arrow, 1931 [species]
Onthophagus pallens D'orbigny, 1908 [species]
Onthophagus pallidipennis Fahraeus, 1857 [species]
Onthophagus pallidus D'orbigny, 1905 [species]
Onthophagus paluma Matthews, 1972 [species]
Onthophagus panfilovi Kabakov, 1983 [species]
Onthophagus panici Petrovitz, 1964 [species]
Onthophagus panoplus Bates, 1888 [species]
Onthophagus paolae Palestrini, 1992 [species]
Onthophagus papuensis Harold, 1877 [species]
Onthophagus papulatorius Kabakov, 2008 [species]
Onthophagus papulatus Boucomont, 1914 [species]
Onthophagus parachandrai Ochi, Kon & Tsubaki, 2009 [species]
Onthophagus parafalculatus Ochi, Kon & Tsubaki, 2009 [species]
Onthophagus parafasciatus Balthasar, 1974 [species]
Onthophagus parallelicornis Macleay, 1887 [species]
Onthophagus paramasaoi Ochi, Kon & Barclay, 2009 [species]
Onthophagus parapalatus KRIKKEN & HUIJBREGTS, 1988 [species]
Onthophagus parapedisequus Balthasar, 1969 [species]
Onthophagus paravinctus Tagliaferri & Moretto, 2012 [species]
Onthophagus parceguttatus D'orbigny, 1902 [species]
Onthophagus parcenotatus D'orbigny, 1913 [species]
Onthophagus parcepictus D'orbigny, 1908 [species]
Onthophagus parcepilosus Balthasar, 1966 [species]
Onthophagus pardalis Fabricius, 1798 [species]
Onthophagus parenthesis Boucomont, 1913 [species]
Onthophagus parisii Balthasar, 1941 [species]
Onthophagus parmatus Reitter, 1892 [species]
Onthophagus paroculus Krikken & Huijbregts, 1987 [species]
Onthophagus parrumbal Matthews, 1972 [species]
Onthophagus parryi Harold, 1869 [species]
Onthophagus parumnotatus Fahraeus, 1857 [species]
Onthophagus parvidens Frey, 1971 [species]
Onthophagus parvifrons D'orbigny, 1907 [species]
Onthophagus parviobscurior Ochi, Kon & Tsubaki, 2009 [species]
Onthophagus parvulus Fabricius, 1798 [species]
Onthophagus parvus Blanchard, 1853 [species]
Onthophagus pastillatus Boucomont, 1919 [species]
Onthophagus patinatus Marcus, 1917 [species]
Onthophagus paucigranosus Lansberge, 1886 [species]
Onthophagus paulianellus Frey, 1958 [species]
Onthophagus pauliani Balthasar, 1937 [species]
Onthophagus pauper Boucomont, 1914 [species]
Onthophagus pauxillus D'orbigny, 1902 [species]
Onthophagus pavidus Harold, 1877 [species]
Onthophagus pedator Sharp, 1875 [species]
Onthophagus pedester Howden & Génier, 2004 [species]
Onthophagus pedisequus Balthasar, 1969 [species]
Onthophagus pegesimallus Marcus, 1917 [species]
Onthophagus pendjarius Josso & Prévost, 2006 [species]
Onthophagus penicillatus Harold, 1879 [species]
Onthophagus peninsularis Boucomont, 1914 [species]
Onthophagus peninsulocupreus Krikken & Huijbregts, 2009 [species]
Onthophagus peninsulomerus Huijbregts & Krikken, 2011 [species]
Onthophagus penmani Masumoto & Ochi & Hanboonsong, 2002 [species]
Onthophagus pentacanthus Harold, 1867 [species]
Onthophagus peramelinus Lea, 1923 [species]
Onthophagus perniger Boucomont, 1930 [species]
Onthophagus perpilosus Macleay, 1871 [species]
Onthophagus personatus Boucomont, 1914 [species]
Onthophagus petenensis HOWDEN & GILL, 1993 [species]
Onthophagus petrovitzi Frey, 1975 [species]
Onthophagus petrovitzianus Pittino, 2006 [species]
Onthophagus pexatus Harold, 1869 [species]
Onthophagus peyrierasi Paulian, 1985 [species]
Onthophagus phanaeicollis Lansberge, 1883 [species]
Onthophagus phanaeides Frey, 1956 [species]
Onthophagus phanaeiformis Boucomont, 1914 [species]
Onthophagus phanaeomorphus Janssens, 1954 [species]
Onthophagus phatoensis Masumoto, Ochi & Hanboonsong, 2008 [species]
Onthophagus phetchabunensis Masumoto, Ochi & Hanboonsong, 2002 [species]
Onthophagus philippinensis Boucomont, 1919 [species]
Onthophagus phillippsorum Krikken & Huijbregts, 1987 [species]
Onthophagus phoenicocerus Lea, 1923 [species]
Onthophagus phraoensis Masumoto, 1990 [species]
Onthophagus phrixus Balthasar, 1969 [species]
Onthophagus phrutsaphaakhomus Masumoto, 1992 [species]
Onthophagus phukhieoensis Masumoto, Ochi & Hanboonsong, 2007 [species]
Onthophagus phuquoci Paulian, 1945 [species]
Onthophagus picatus D'orbigny, 1902 [species]
Onthophagus piceiceps D'orbigny, 1904 [species]
Onthophagus piceorufulus Kabakov, 1994 [species]
Onthophagus picipennis Hope, 1841 [species]
Onthophagus pictipennis D'orbigny, 1913 [species]
Onthophagus pictipodex D'orbigny, 1902 [species]
Onthophagus picturatus D'orbigny, 1908 [species]
Onthophagus pictus Reitter, 1893 [species]
Onthophagus piffli Petrovitz, 1961 [species]
Onthophagus pilicollis D'orbigny, 1902 [species]
Onthophagus pilipodex D'orbigny, 1913 [species]
Onthophagus pillara Matthews, 1972 [species]
Onthophagus pilosus Fahraeus, 1857 [species]
Onthophagus pilularius Lansberge, 1883 [species]
Onthophagus pimpasaleei Masumoto, Ochi & Hanboonsong, 2008 [species]
Onthophagus pinaroo STOREY & WEIR, 1990 [species]
Onthophagus pinguis Gerstaecker, 1871 [species]
Onthophagus pipitzi Ancey, 1883 [species]
Onthophagus pisciphagus WALTER & CAMBEFORT, 1977 [species]
Onthophagus piyawati Masumoto, Ochi & Hanboonsong, 2002 [species]
Onthophagus placens Péringuey, 1904 [species]
Onthophagus planaticeps D'orbigny, 1913 [species]
Onthophagus planiceps Macleay, 1886 [species]
Onthophagus planicollis Harold, 1880 [species]
Onthophagus planifrons Frey, 1962 [species]
Onthophagus platalea Arrow, 1941 [species]
Onthophagus plato Bates, 1888 [species]
Onthophagus platypus Zhang, 1997 [species]
Onthophagus plebejus Klug, 1855 [species]
Onthophagus pleurogonus D'orbigny, 1913 [species]
Onthophagus plicatifrons D'orbigny, 1908 [species]
Onthophagus poeophagus Kabakov, 1979 [species]
Onthophagus poggii Scheuern, 1996 [species]
Onthophagus politissimus D'orbigny, 1908 [species]
Onthophagus politus Fabricius, 1798 [species]
Onthophagus pollicatus Harold, 1879 [species]
Onthophagus polyedrus D'orbigny, 1905 [species]
Onthophagus polyodon D'orbigny, 1913 [species]
Onthophagus polystigma D'orbigny, 1902 [species]
Onthophagus ponticus Harold, 1883 [species]
Onthophagus pooensis Cambefort & Nicolas, 1991 [species]
Onthophagus popovi Kabakov, 1983 [species]
Onthophagus porcus Arrow, 1931 [species]
Onthophagus poringensis Ochi & Kon, 2005 [species]
Onthophagus porrectus Reiche, 1847 [species]
Onthophagus possoi Walter, 1982 [species]
Onthophagus posticicornis D'orbigny, 1913 [species]
Onthophagus posticus Erichson, 1842 [species]
Onthophagus praecavatus D'Orbigny, 1913 [species]
Onthophagus praecellens Bates, 1887 [species]
Onthophagus praedatus Harold, 1862 [species]
Onthophagus praedentatus D'Orbigny, 1904 [species]
Onthophagus praefossus D'Orbigny, 1913 [species]
Onthophagus praelaminatus Frey, 1957 [species]
Onthophagus praestans Péringuey, 1901 [species]
Onthophagus praetortus Péringuey, 1904 [species]
Onthophagus prehensilis Arrow, 1920 [species]
Onthophagus privus Kabakov, 1998 [species]
Onthophagus probus Péringuey, 1901 [species]
Onthophagus procurvus Balthasar, 1935 [species]
Onthophagus producticollis d'Orbigny, 1908 [species]
Onthophagus productus Arrow, 1907 [species]
Onthophagus profanus Frey, 1958 [species]
Onthophagus proletarius Harold, 1875 [species]
Onthophagus promontorii Boucomont, 1924 [species]
Onthophagus pronus Erichson, 1842 [species]
Onthophagus propinquus Macleay, 1888 [species]
Onthophagus propraecellens HOWDEN & GILL, 1987 [species]
Onthophagus prostans Reiche, 1847 [species]
Onthophagus proteus D'orbigny, 1902 [species]
Onthophagus protuberans Frey, 1955 [species]
Onthophagus proximus D'orbigny, 1908 [species]
Onthophagus pseudoaeneus D'orbigny, 1908 [species]
Onthophagus pseudoalcyon d'Orbigny, 1913 [species]
Onthophagus pseudoarmatus Balthasar, 1944 [species]
Onthophagus pseudobidens D'orbigny, 1915 [species]
Onthophagus pseudobrutus Kabakov, 1983 [species]
Onthophagus pseudocaccobius Reitter, 1889 [species]
Onthophagus pseudocoracinus Kabakov, 1983 [species]
Onthophagus pseudofimetarius Balthasar, 1946 [species]
Onthophagus pseudoflexicollis Moretto, 2004 [species]
Onthophagus pseudohystrix Masumoto, 1995 [species]
Onthophagus pseudojavanus Paulian, 1931 [species]
Onthophagus pseudoliberianus Moretto, 2010 [species]
Onthophagus pseudopilosus Frey, 1958 [species]
Onthophagus pseudoplebejus Moretto, 2009 [species]
Onthophagus pseudorudis Ochi & Araya, 1992 [species]
Onthophagus pseudosanguineus WALTER & CAMBEFORT, 1977 [species]
Onthophagus pseudosellatus Balthasar, 1964 [species]
Onthophagus pseudovinctus Tagliaferri & Moretto, 2012 [species]
Onthophagus pseudovirens D'Orbigny, 1913 [species]
Onthophagus pseudoworoae Ochi, Kon & Barclay, 2009 [species]
Onthophagus psychopompus Ziani & Garakhloo, 2010 [species]
Onthophagus ptox Erichson, 1847 [species]
Onthophagus puberulus D'orbigny, 1902 [species]
Onthophagus pugionatus Fahraeus, 1857 [species]
Onthophagus pugnacior Blackburn, 1903 [species]
Onthophagus pugnax Harold, 1868 [species]
Onthophagus pulchellus D'orbigny, 1905 [species]
Onthophagus pullatus D'orbigny, 1905 [species]
Onthophagus pullus Roth, 1851 [species]
Onthophagus punctator Reitter, 1893 [species]
Onthophagus punctatus Illiger, 1803 [species]
Onthophagus punneeae Masumoto, 1989 [species]
Onthophagus punthari Storey, 1977 [species]
Onthophagus pupillatus Kolbe, 1886 [species]
Onthophagus purifrons D'orbigny, 1908 [species]
Onthophagus purpurascens Boucomont, 1914 [species]
Onthophagus purpureicollis Macleay, 1864 [species]
Onthophagus pusillus Fabricius, 1798 [species]
Onthophagus pusio Fahraeus, 1857 [species]
Onthophagus pygargus Motschulsky, 1845 [species]
Onthophagus pygidialis Lansberge, 1883 [species]
Onthophagus pygmaeus Schaller, 1783 [species]
Onthophagus pyramidalis Klug, 1855 [species]
Onthophagus quadraticeps Harold, 1867 [species]
Onthophagus quadriarmatus Fairmaire, 1892 [species]
Onthophagus quadricallosus D'orbigny, 1902 [species]
Onthophagus quadricolor Kabakov, 1983 [species]
Onthophagus quadricristatus D'Orbigny, 1908 [species]
Onthophagus quadricuspis d'Orbigny, 1908 [species]
Onthophagus quadridentatus Fabricius, 1798 [species]
Onthophagus quadrilunatus D'orbigny, 1902 [species]
Onthophagus quadrimaculatus Raffray, 1877 [species]
Onthophagus quadrinodosus Fahraeus, 1857 [species]
Onthophagus quadrinodus Reitter, 1896 [species]
Onthophagus quadrinotatus D'orbigny, 1905 [species]
Onthophagus quadripustulatus Fabricius, 1775 [species]
Onthophagus quadrispinosus D'Orbigny, 1908 [species]
Onthophagus quadrituber D'Orbigny, 1908 [species]
Onthophagus quaestus Sharp, 1875 [species]
Onthophagus quasijohkii Ochi & Kon, 2005 [species]
Onthophagus quasitagal Ochi & Kon, 2005 [species]
Onthophagus queenslandicus Blackburn, 1903 [species]
Onthophagus quetzalis HOWDEN & GILL, 1993 [species]
Onthophagus quezonensis Ochi & Araya, 1992 [species]
Onthophagus quinquetuberculatus Macleay, 1871 [species]
Onthophagus quiproquo Moretto & Génier, 2010 [species]
Onthophagus rachelis Martin-Piera, 1985 [species]
Onthophagus ragazzii D'orbigny, 1904 [species]
Onthophagus rakovici Pittino, 2004 [species]
Onthophagus ramosellus Bates, 1891 [species]
Onthophagus ramosicornis d'Orbigny, 1902 [species]
Onthophagus ramosus Wiedemann, 1823 [species]
Onthophagus rana Arrow, 1931 [species]
Onthophagus rangifer Klug, 1855 [species]
Onthophagus ranongensis Masumoto, Ochi & Hanboonsong, 2008 [species]
Onthophagus ranunculus Arrow, 1913 [species]
Onthophagus rarus Guérin-Méneville, 1829 [species]
Onthophagus rasidorsis D'Orbigny, 1905 [species]
Onthophagus rasipennis D'orbigny, 1908 [species]
Onthophagus ratchasimaensis Masumoto & Hanboonsong & Ochi, 2002 [species]
Onthophagus rechingeri Petrovitz, 1980 [species]
Onthophagus rectecornutus Lansberge, 1883 [species]
Onthophagus rectefurcatus Fairmaire, 1891 [species]
Onthophagus rectestriatus D'orbigny, 1915 [species]
Onthophagus rectilamina D'orbigny, 1902 [species]
Onthophagus rectorispauliani Cambefort, 1980 [species]
Onthophagus refulgens Arrow, 1931 [species]
Onthophagus regalis Arrow, 1907 [species]
Onthophagus remotus Harold, 1879 [species]
Onthophagus renaudpauliani Ochi & Araya, 1996 [species]
Onthophagus reticollis Macleay, 1886 [species]
Onthophagus reticulatus D'orbigny, 1902 [species]
Onthophagus reticuliger Frey, 1960 [species]
Onthophagus revoili Lansberge, 1882 [species]
Onthophagus reyesi ZUNINO & HALFFTER, 1988 [species]
Onthophagus rhinocerus Gomes Alves, 1944 [species]
Onthophagus rhinolophus Harold, 1869 [species]
Onthophagus rhinophyllus Harold, 1868 [species]
Onthophagus rhodesianus Frey, 1975 [species]
Onthophagus rhombocephalus Kabakov, 1983 [species]
Onthophagus rhynchophorus Péringuey, 1904 [species]
Onthophagus ribbei Boucomont, 1914 [species]
Onthophagus riekoae Ochi & Kon, 2005 [species]
Onthophagus riparius Lansberge, 1885 [species]
Onthophagus ritsemai Lansberge, 1883 [species]
Onthophagus robertopoggii Ochi & Kon, 2006 [species]
Onthophagus rodentium Pittino, 2004 [species]
Onthophagus rorarius Harold, 1877 [species]
Onthophagus rosettae Frey, 1958 [species]
Onthophagus rostratus Harold, 1869 [species]
Onthophagus rotundatus D'orbigny, 1905 [species]
Onthophagus rotundibasis D'orbigny, 1902 [species]
Onthophagus rotundicollis Lansberge, 1883 [species]
Onthophagus roubali Balthasar, 1935 [species]
Onthophagus rougonorum Cambefort, 1984 [species]
Onthophagus rouyeri Boucomont, 1914 [species]
Onthophagus royi Biswas & Chatterjee, 1985 [species]
Onthophagus ruandanus Frey, 1971 [species]
Onthophagus rubefactus D'orbigny, 1913 [species]
Onthophagus rubellus D'orbigny, 1908 [species]
Onthophagus rubens D'orbigny, 1902 [species]
Onthophagus rubenticollis D'orbigny, 1902 [species]
Onthophagus rubescens Macleay, 1888 [species]
Onthophagus rubicundulus Macleay, 1871 [species]
Onthophagus rubidus D'orbigny, 1913 [species]
Onthophagus rubrescens Blanchard, 1843 [species]
Onthophagus rubricatus D'orbigny, 1902 [species]
Onthophagus rubricollis Hope, 1831 [species]
Onthophagus rubrimaculatus Macleay, 1864 [species]
Onthophagus rubripennis Arrow, 1907 [species]
Onthophagus rudis Sharp, 1875 [species]
Onthophagus rufescens Bates, 1887 [species]
Onthophagus ruficapillus Brullé, 1832 [species]
Onthophagus ruficauda Arrow, 1931 [species]
Onthophagus rufimanus Kabakov, 1982 [species]
Onthophagus rufiobscurior Ochi, Kon & Tsubaki, 2009 [species]
Onthophagus rufipodex D'Orbigny, 1913 [species]
Onthophagus rufobasalis Fairmaire, 1887 [species]
Onthophagus rufocastaneus Fairmaire, 1887 [species]
Onthophagus rufolimbatus D'orbigny, 1913 [species]
Onthophagus rufonotatus D'orbigny, 1902 [species]
Onthophagus rufopygus Frey, 1957 [species]
Onthophagus rufosignatus Macleay, 1864 [species]
Onthophagus rufostillans D'orbigny, 1907 [species]
Onthophagus rufovirens D'orbigny, 1904 [species]
Onthophagus rugicollis Harold, 1880 [species]
Onthophagus rugidorsis D'orbigny, 1913 [species]
Onthophagus rugipennis D'orbigny, 1908 [species]
Onthophagus rugosipennis Frey, 1973 [species]
Onthophagus rugosissimus D'orbigny, 1913 [species]
Onthophagus rugulipennis Fairmaire, 1887 [species]
Onthophagus rugulosus Harold, 1886 [species]
Onthophagus rupicapra Waterhouse, 1894 [species]
Onthophagus sabahensis Ochi & Kon, 2006 [species]
Onthophagus sacharovskii Olsoufieff, 1918 [species]
Onthophagus saggitarius Fabricius, 1775 [species]
Onthophagus sagittarius Fabricius, 1775 [species]
Onthophagus sahai Biswas & Chatterjee, 1986 [species]
Onthophagus saïgonensis Boucomont, 1924 [species]
Onthophagus sakaeratensis Masumoto & Hanboonsong & Ochi, 2002 [species]
Onthophagus sakainoi Masumoto, 1991 [species]
Onthophagus sakamakii Ochi, Kon & Kawahara, 2010 [species]
Onthophagus salakensis Ochi, Kon & Hartini, 2010 [species]
Onthophagus salebrosus Macleay, 1888 [species]
Onthophagus saleyeri Lansberge, 1883 [species]
Onthophagus salvadorensis ZUNINO & HALFFTER, 1988 [species]
Onthophagus salvazai Paulian, 1978 [species]
Onthophagus samai Ziani, 2011 [species]
Onthophagus samoengus Masumoto, Ochi & Hanboonsong, 2008 [species]
Onthophagus sanggona Huijbregts & Krikken, 2009 [species]
Onthophagus sangirensis Boucomont, 1914 [species]
Onthophagus sanguineus D'orbigny, 1902 [species]
Onthophagus sanguinolentus D'orbigny, 1908 [species]
Onthophagus sangwalus Masumoto, Ochi & Hanboonsong, 2008 [species]
Onthophagus sansibaricus Harold, 1878 [species]
Onthophagus sapphirinus Fahraeus, 1857 [species]
Onthophagus sarasinorum Krikken & Huijbregts, 2011 [species]
Onthophagus sarawacus Harold, 1877 [species]
Onthophagus sasajii Ochi & Kon, 2001 [species]
Onthophagus satoi Ochi, Shimada & Kon, 2003 [species]
Onthophagus saudiensis Frey, 1962 [species]
Onthophagus sauteri Gillet, 1924 [species]
Onthophagus savanicola Cambefort, 1984 [species]
Onthophagus sayapensis Ochi & Kon, 2005 [species]
Onthophagus scaber Roth, 1851 [species]
Onthophagus scaberrimus D'orbigny, 1904 [species]
Onthophagus scabriusculus Harold, 1873 [species]
Onthophagus scapularis D'orbigny, 1902 [species]
Onthophagus sceptrifer Boucomont, 1924 [species]
Onthophagus schaefernai Balthasar, 1935 [species]
Onthophagus schaufussi Harold, 1867 [species]
Onthophagus schawalleri Scheuern, 1996 [species]
Onthophagus schillhammeri Kabakov, 2006 [species]
Onthophagus schoolmeestersi Ochi & Kon, 2007 [species]
Onthophagus schunckei Paulian, 1936 [species]
Onthophagus schwaneri Snellen van Vollenhoven, 1864 [species]
Onthophagus sciron Balthasar, 1969 [species]
Onthophagus scotti Masumoto, Ochi & Hanboonsong, 2002 [species]
Onthophagus scrutator Harold, 1877 [species]
Onthophagus sculptilis Gerstaecker, 1871 [species]
Onthophagus seabrai Gomes Alves, 1944 [species]
Onthophagus secundarius Roth, 1851 [species]
Onthophagus sellatulus D'orbigny, 1902 [species]
Onthophagus sellatus Klug, 1845 [species]
Onthophagus sembeli Krikken & Huijbregts, 2008 [species]
Onthophagus semiaratus D'orbigny, 1902 [species]
Onthophagus semiasper D'orbigny, 1902 [species]
Onthophagus semiaureus Lansberge, 1883 [species]
Onthophagus semichalcites D'orbigny, 1902 [species]
Onthophagus semicínctus D'Orbigny, 1897 [species]
Onthophagus semicornis Panzer, 1798 [species]
Onthophagus semicroceus D'orbigny, 1915 [species]
Onthophagus semicupreus Harold, 1877 [species]
Onthophagus semidanumensis Ochi, Kon & Barclay, 2009 [species]
Onthophagus semifex Krikken & Huijbregts, 2008 [species]
Onthophagus semiflavus Boheman, 1860 [species]
Onthophagus semigraniger D'orbigny, 1905 [species]
Onthophagus semigranosus Lansberge, 1883 [species]
Onthophagus semiiris Thomson, 1858 [species]
Onthophagus semilaevis D'orbigny, 1913 [species]
Onthophagus semimetallicus Lea, 1923 [species]
Onthophagus seminitens D'orbigny, 1902 [species]
Onthophagus seminitidus D'orbigny, 1908 [species]
Onthophagus semiopacus Harold, 1869 [species]
Onthophagus semipacificus Ochi & Kon, 2006 [species]
Onthophagus semipersonatus Ochi, Kon & Tsubaki, 2009 [species]
Onthophagus semipiceus Kabakov, 1998 [species]
Onthophagus semivestitus Paulian, 1937 [species]
Onthophagus semivirescens D'orbigny, 1902 [species]
Onthophagus semiviridis D'orbigny, 1904 [species]
Onthophagus senegalensis D'orbigny, 1902 [species]
Onthophagus senescens Péringuey, 1908 [species]
Onthophagus senex Boucomont, 1914 [species]
Onthophagus seniculus Fabricius, 1781 [species]
Onthophagus sepilokensis Ochi & Kon, 2006 [species]
Onthophagus seramicus Huijbregts & Krikken, 2012 [species]
Onthophagus serapiensis Ochi, Kon & Barclay, 2008 [species]
Onthophagus serdangensis Lansberge, 1886 [species]
Onthophagus sericans Frey, 1975 [species]
Onthophagus sericatus Reitter, 1893 [species]
Onthophagus sericeicollis Boucomont, 1928 [species]
Onthophagus serienotatus D'orbigny, 1902 [species]
Onthophagus setchan Masumoto, 1984 [species]
Onthophagus setoculus Krikken & Huijbregts, 1987 [species]
Onthophagus setosus Fahraeus, 1857 [species]
Onthophagus sexcornutus d'Orbigny, 1902 [species]
Onthophagus sexdentatus Boucomont, 1919 [species]
Onthophagus sexstriatus Montrouzier, 1855 [species]
Onthophagus shanicus Kabakov, 2006 [species]
Onthophagus shapovalovi Gusakov, 2012 [species]
Onthophagus sharpi Harold, 1875 [species]
Onthophagus shibatai Nakane, 1960 [species]
Onthophagus shillongensis Scheuern, 1995 [species]
Onthophagus shimba Cambefort, 1980 [species]
Onthophagus shirakii Nakane, 1960 [species]
Onthophagus shizumui Ochi & Araya, 1992 [species]
Onthophagus sibela Huijbregts & Krikken, 2012 [species]
Onthophagus sibiricus Harold, 1877 [species]
Onthophagus sibuyanus Boucomont, 1924 [species]
Onthophagus sidama Gestro, 1895 [species]
Onthophagus sideki Krikken & Huijbregts, 1987 [species]
Onthophagus signaticollis Frey, 1970 [species]
Onthophagus signatus Fahraeus, 1857 [species]
Onthophagus signifer Harold, 1877 [species]
Onthophagus sihkahonoi Huijbregts & Krikken, 2009 [species]
Onthophagus sikkimensis Gillet, 1925 [species]
Onthophagus silus Balthasar, 1960 [species]
Onthophagus simboroni Ochi & Kon, 2006 [species]
Onthophagus similis Scriba, 1790 [species]
Onthophagus simillimus D'orbigny, 1913 [species]
Onthophagus simius Reitter, 1893 [species]
Onthophagus simoni D'orbigny, 1902 [species]
Onthophagus simplex Raffray, 1877 [species]
Onthophagus simpliciceps D'orbigny, 1905 [species]
Onthophagus simplicifrons Reitter, 1893 [species]
Onthophagus simulator D'orbigny, 1905 [species]
Onthophagus sinagai Krikken & Huijbregts, 2008 [species]
Onthophagus singhaakhomus Masumoto, 1992 [species]
Onthophagus singulariformis Kohlmann & Solis, 2001 [species]
Onthophagus sinicus Zhang & Wang, 1997 [species]
Onthophagus sinuosicollis D'orbigny, 1905 [species]
Onthophagus sinuosus D'orbigny, 1913 [species]
Onthophagus sipilouensis Cambefort, 1984 [species]
Onthophagus sisyphoides Krikken, 1977 [species]
Onthophagus sjoestethi D'Orbigny, 1904 [species]
Onthophagus sloanei Blackburn, 1903 [species]
Onthophagus smeenki Cambefort, 1980 [species]
Onthophagus smetanai Balthasar, 1952 [species]
Onthophagus snoflaki Balthasar, 1944 [species]
Onthophagus sobrius Balthasar, 1959 [species]
Onthophagus socialis Arrow, 1931 [species]
Onthophagus solidus Gillet, 1927 [species]
Onthophagus solisi HOWDEN & GILL, 1993 [species]
Onthophagus solivagus Harold, 1886 [species]
Onthophagus solmani Stebnicka, 1975 [species]
Onthophagus somalicola Balthasar, 1941 [species]
Onthophagus somalicus d'Orbigny, 1902 [species]
Onthophagus sonani Miwa, 1930 [species]
Onthophagus songsokensis Biswas & Chatterjee, 1985 [species]
Onthophagus sophiae Nicolas, 2006 [species]
Onthophagus sparsepunctatus Frey, 1956 [species]
Onthophagus sparsulus Reitter, 1893 [species]
Onthophagus spathatus Boucomont, 1914 [species]
Onthophagus speculatus D'Orbigny, 1908 [species]
Onthophagus speculicollis Quedenfeldt, 1884 [species]
Onthophagus speculifer Solsky, 1876 [species]
Onthophagus spiculatus Boucomont, 1914 [species]
Onthophagus spinicornis Gillet, 1930 [species]
Onthophagus spinifex Fabricius, 1781 [species]
Onthophagus splendidus Boucomont, 1932 [species]
Onthophagus spurcatus D'orbigny, 1905 [species]
Onthophagus squalidus Lea, 1923 [species]
Onthophagus stanleyi Moretto & Génier, 2010 [species]
Onthophagus stehliki Balthasar, 1966 [species]
Onthophagus steinheili Harold, 1880 [species]
Onthophagus stellio Erichson, 1843 [species]
Onthophagus stenocerus Harold, 1867 [species]
Onthophagus sternalis Arrow, 1931 [species]
Onthophagus sternax Balthasar, 1959 [species]
Onthophagus sticticus Harold, 1867 [species]
Onthophagus stictus D'orbigny, 1913 [species]
Onthophagus stigmosus D'orbigny, 1902 [species]
Onthophagus stillatus D'orbigny, 1904 [species]
Onthophagus stockwelli Howden & Young, 1981 [species]
Onthophagus stomachosus Krell, 2009 [species]
Onthophagus strabo Reitter, 1892 [species]
Onthophagus strandi Balthasar, 1935 [species]
Onthophagus streltsovi Tarasov & Kabakov, 2010 [species]
Onthophagus striaticollis Kabakov, 1983 [species]
Onthophagus strictestriatus D'orbigny, 1913 [species]
Onthophagus strnadi Kabakov, 1998 [species]
Onthophagus stuhlmanni D'orbigny, 1908 [species]
Onthophagus stylocerus Graells, 1851 [species]
Onthophagus subalternans D'orbigny, 1902 [species]
Onthophagus subansiriensis Biswas, 1979 [species]
Onthophagus subcancer Howden, 1973 [species]
Onthophagus subcinctus D'orbigny, 1913 [species]
Onthophagus subcornutus Boucomont, 1914 [species]
Onthophagus subdivisus D'orbigny, 1908 [species]
Onthophagus subextensus Kolbe, 1895 [species]
Onthophagus subglaber D'orbigny, 1902 [species]
Onthophagus subhumeralis D'orbigny, 1902 [species]
Onthophagus subnudus D'orbigny, 1902 [species]
Onthophagus subocellatus D'orbigny, 1905 [species]
Onthophagus subplanus D'orbigny, 1902 [species]
Onthophagus subrugosus D'orbigny, 1905 [species]
Onthophagus subsapaensis Kabakov, 1994 [species]
Onthophagus subsulcatus D'orbigny, 1908 [species]
Onthophagus subulifer D'orbigny, 1908 [species]
Onthophagus suermelii Petrovitz, 1963 [species]
Onthophagus suffusus Klug, 1855 [species]
Onthophagus sugihartoi Ochi, 2007 [species]
Onthophagus sugillatus Klug, 1855 [species]
Onthophagus suginoi Ochi, 1984 [species]
Onthophagus suginokoichii Ochi & Kon, 2008 [species]
Onthophagus suillus Arrow, 1931 [species]
Onthophagus sulawesiensis Krikken & Huijbregts, 2011 [species]
Onthophagus sulcatulus D'orbigny, 1907 [species]
Onthophagus sulci Balthasar, 1935 [species]
Onthophagus sulcipennis D'orbigny, 1902 [species]
Onthophagus sumatramontanus Ochi & Kon, 2008 [species]
Onthophagus sumatranus Lansberge, 1883 [species]
Onthophagus sumawacus Krikken & Huijbregts, 2008 [species]
Onthophagus sumbavensis Boucomont, 1914 [species]
Onthophagus sumptuosus Paulian, 1937 [species]
Onthophagus sunantaae Masumoto, 1989 [species]
Onthophagus sundanensis Lansberge, 1883 [species]
Onthophagus superbus D'Orbigny, 1904 [species]
Onthophagus surdus Boucomont, 1925 [species]
Onthophagus susterai Balthasar, 1952 [species]
Onthophagus sutiliceps D'Orbigny, 1902 [species]
Onthophagus sutleinensis Splichal, 1910 [species]
Onthophagus suturalis Péringuey, 1888 [species]
Onthophagus suturellus Brullé, 1832 [species]
Onthophagus sycophanta Fairmaire, 1887 [species]
Onthophagus sydneyensis Blackburn, 1903 [species]
Onthophagus sylvestris WALTER & CAMBEFORT, 1977 [species]
Onthophagus symbioticus Arrow, 1920 [species]
Onthophagus synceri Cambefort, 1984 [species]
Onthophagus taayai Masumoto, 1995 [species]
Onthophagus tabellicornis Macleay, 1864 [species]
Onthophagus tabellifer Gillet, 1927 [species]
Onthophagus tabidus Balthasar, 1935 [species]
Onthophagus taboranus D'orbigny, 1902 [species]
Onthophagus taeniatus Boucomont, 1914 [species]
Onthophagus tagal Boucomont, 1924 [species]
Onthophagus taichii Ochi, Kon & Barclay, 2008 [species]
Onthophagus taiensis Cambefort, 1984 [species]
Onthophagus taiwanus Nomura, 1973 [species]
Onthophagus taiyaruensis Masumoto, 1977 [species]
Onthophagus takakui Ochi, Kon & Hartini, 2009 [species]
Onthophagus takedai Ochi & Araya, 1992 [species]
Onthophagus takeshii Ochi & Araya, 1992 [species]
Onthophagus talpa Fahraeus, 1857 [species]
Onthophagus tambing Krikken & Huijbregts, 2011 [species]
Onthophagus tamijii Kon, Sakai & Ochi, 2000 [species]
Onthophagus tamworthi Blackburn, 1903 [species]
Onthophagus tanganus D'orbigny, 1913 [species]
Onthophagus taoi Ochi & Kon, 2006 [species]
Onthophagus tapirus Sharp, 1877 [species]
Onthophagus taprobanus Arrow, 1931 [species]
Onthophagus tarandus Fabricius, 1792 [species]
Onthophagus tarsius Arrow, 1941 [species]
Onthophagus taruni Biswas & Chatterjee, 1986 [species]
Onthophagus tatsienluensis Balthasar, 1942 [species]
Onthophagus taurinus White, 1844 [species]
Onthophagus tauroides Gillet, 1930 [species]
Onthophagus taxillus Balthasar, 1969 [species]
Onthophagus taymansi D'Orbigny, 1908 [species]
Onthophagus teitanicus D'orbigny, 1902 [species]
Onthophagus telegonus Balthasar, 1969 [species]
Onthophagus telephus Balthasar, 1969 [species]
Onthophagus temporalis D'orbigny, 1902 [species]
Onthophagus tenax Balthasar, 1964 [species]
Onthophagus tenebrosus Harold, 1871 [species]
Onthophagus tenuigraniger D'orbigny, 1913 [species]
Onthophagus tenuistriatus D'orbigny, 1905 [species]
Onthophagus terminatus Eschscholtz, 1822 [species]
Onthophagus terrara Storey, 1977 [species]
Onthophagus tersicollis Müller, 1947 [species]
Onthophagus tersidorsis D'Orbigny, 1902 [species]
Onthophagus tersipennis D'orbigny, 1902 [species]
Onthophagus tersus D'orbigny, 1913 [species]
Onthophagus tesquorum Semenov & Medvedev, 1927 [species]
Onthophagus tesseratus D'orbigny, 1908 [species]
Onthophagus tessulatus Harold, 1871 [species]
Onthophagus testaceoviolaceus Paulian, 1937 [species]
Onthophagus tetricus Harold, 1877 [species]
Onthophagus thai Kabakov, 1994 [species]
Onthophagus thailaevis Masumoto, Ochi & Hanboonsong, 2008 [species]
Onthophagus thainuaensis Masumoto, Ochi & Hanboonsong, 2008 [species]
Onthophagus thaitai Masumoto, Ochi & Hanboonsong, 2008 [species]
Onthophagus thanwaakhomus Masumoto, 1992 [species]
Onthophagus tholaayi Masumoto, 1990 [species]
Onthophagus thoreyi Harold, 1868 [species]
Onthophagus tiamicus Kabakov, 1994 [species]
Onthophagus tibetanus Arrow, 1907 [species]
Onthophagus tigrinus D'orbigny, 1908 [species]
Onthophagus timorensis Boucomont, 1914 [species]
Onthophagus tiniocelloides Boucomont, 1925 [species]
Onthophagus tirapensis Biswas & Chatterjee, 1985 [species]
Onthophagus togeman Matthews, 1972 [species]
Onthophagus tongbantumi Masumoto, Ochi & Hanboonsong, 2002 [species]
Onthophagus tonkineus Gillet, 1921 [species]
Onthophagus tonsus D'orbigny, 1902 [species]
Onthophagus tonywhitteni Krikken & Huijbregts, 2011 [species]
Onthophagus toraut Krikken & Huijbregts, 2011 [species]
Onthophagus totonicapamus Bates, 1887 [species]
Onthophagus traginus Frey, 1972 [species]
Onthophagus tragoides Boucomont, 1914 [species]
Onthophagus tragus Fabricius, 1792 [species]
Onthophagus transisthmius Howden & Young, 1981 [species]
Onthophagus transquadridentatus Scheuern, 1995 [species]
Onthophagus transvestitus Huijbregts & Krikken, 2009 [species]
Onthophagus trapezicornis D'orbigny, 1902 [species]
Onthophagus traversii D'orbigny, 1904 [species]
Onthophagus trawalla STOREY & WEIR, 1990 [species]
Onthophagus triacanthus Castelnau, 1840 [species]
Onthophagus trianguliceps D'Orbigny, 1905 [species]
Onthophagus triarmatus D'Orbigny, 1904 [species]
Onthophagus tricariniger D'orbigny, 1902 [species]
Onthophagus tricavicollis Lea, 1923 [species]
Onthophagus triceratops Arrow, 1913 [species]
Onthophagus trichopygus D'orbigny, 1905 [species]
Onthophagus tricolor Boucomont, 1914 [species]
Onthophagus tricornifrons D'Orbigny, 1908 [species]
Onthophagus tricorniger Boheman, 1860 [species]
Onthophagus tricornis Wiedemann, 1823 [species]
Onthophagus tridenticeps D'orbigny, 1902 [species]
Onthophagus tridentitibialis Ochi & Kon, 2008 [species]
Onthophagus trifidisetis D'orbigny, 1905 [species]
Onthophagus trigibber Reitter, 1893 [species]
Onthophagus triimpressus D'orbigny, 1905 [species]
Onthophagus trinodosus Fahraeus, 1857 [species]
Onthophagus trinominatus Goidanich, 1926 [species]
Onthophagus tripartitus D'orbigny, 1902 [species]
Onthophagus tripolitanus Heyden, 1890 [species]
Onthophagus triptolemus Balthasar, 1969 [species]
Onthophagus trispinus Reitter, 1892 [species]
Onthophagus tristis Harold, 1873 [species]
Onthophagus tritinctus Boucomont, 1914 [species]
Onthophagus triundulatus Balthasar, 1964 [species]
Onthophagus trochilus Arrow, 1931 [species]
Onthophagus troglodyta Wiedemann, 1823 [species]
Onthophagus troniceki Balthasar, 1933 [species]
Onthophagus truncaticornis Schaller, 1783 [species]
Onthophagus trungboensis Kabakov, 1983 [species]
Onthophagus tschadensis Balthasar, 1963 [species]
Onthophagus tschoffeni D'orbigny, 1904 [species]
Onthophagus tshuapae Balthasar, 1964 [species]
Onthophagus tsubakii Ochi & Kon, 2009 [species]
Onthophagus tubericollis Raffray, 1877 [species]
Onthophagus tuckonie Matthews, 1972 [species]
Onthophagus tumami Masumoto & Ochi & Hanboonsong, 2002 [species]
Onthophagus tumidulus Gerstaecker, 1871 [species]
Onthophagus tungkamangensis Masumoto, Ochi & Hanboonsong, 2007 [species]
Onthophagus tungkamungensis Masumoto, Ochi & Hanboonsong, 2008 [species]
Onthophagus turbatus Walker, 1858 [species]
Onthophagus turfanicus Balthasar, 1971 [species]
Onthophagus turgidus Kohlmann & Solis, 2012 [species]
Onthophagus turpidoides Kabakov, 2008 [species]
Onthophagus turpidus Reitter, 1887 [species]
Onthophagus turrbal Matthews, 1972 [species]
Onthophagus tuzetae WALTER & CAMBEFORT, 1977 [species]
Onthophagus tweedensis Blackburn, 1903 [species]
Onthophagus uedai Ochi & Kon, 2006 [species]
Onthophagus uelensis Frey, 1960 [species]
Onthophagus ugandicus Frey, 1975 [species]
Onthophagus ukerewensis Balthasar, 1937 [species]
Onthophagus uluguru Frey, 1975 [species]
Onthophagus ulula Balthasar, 1966 [species]
Onthophagus umbilicatus D'orbigny, 1908 [species]
Onthophagus umbilicopunctatus Medvedev, 1959 [species]
Onthophagus umbratus D'orbigny, 1902 [species]
Onthophagus undaticeps D'orbigny, 1902 [species]
Onthophagus unguiculatus Kabakov, 1994 [species]
Onthophagus unicarina D'orbigny, 1904 [species]
Onthophagus unidens Quedenfeldt, 1884 [species]
Onthophagus unidentatus Boucomont, 1927 [species]
Onthophagus unifasciatus Schaller, 1783 [species]
Onthophagus uniformis Heyden, 1886 [species]
Onthophagus urellus Boucomont, 1919 [species]
Onthophagus ursinus D'orbigny, 1902 [species]
Onthophagus ursus Boucomont, 1926 [species]
Onthophagus usambaricus Paulian, 1937 [species]
Onthophagus usurpator Balthasar, 1959 [species]
Onthophagus utsunomiyae Masumoto, Ochi & Hanboonsong, 2007 [species]
Onthophagus vacca Linnaeus, 1767 [species]
Onthophagus vanasseni Krikken & Huijbregts, 2008 [species]
Onthophagus vanderblomi Krikken & Huijbregts, 2008 [species]
Onthophagus vaneyeni Frey, 1960 [species]
Onthophagus vanofwegeni Krikken & Huijbregts, 2008 [species]
Onthophagus varianus Lea, 1923 [species]
Onthophagus variatus D'orbigny, 1902 [species]
Onthophagus variegatus Fabricius, 1798 [species]
Onthophagus variegranosus D'orbigny, 1905 [species]
Onthophagus variolaris Lansberge, 1883 [species]
Onthophagus variolicollis Lea, 1923 [species]
Onthophagus variolosus D'orbigny, 1902 [species]
Onthophagus varius D'orbigny, 1913 [species]
Onthophagus vasiljevi Kabakov, 1983 [species]
Onthophagus vassei D'orbigny, 1908 [species]
Onthophagus vatovai Müller, 1942 [species]
Onthophagus vaulogeri Boucomont, 1923 [species]
Onthophagus velliger D'orbigny, 1907 [species]
Onthophagus ventralis Lansberge, 1883 [species]
Onthophagus ventrosus D'orbigny, 1905 [species]
Onthophagus venzoi Balthasar, 1941 [species]
Onthophagus veracruzensis Delgado & Pensado, 1998 [species]
Onthophagus vermiculatus Frey, 1970 [species]
Onthophagus verrucosus D'orbigny, 1902 [species]
Onthophagus versus D'Orbigny, 1913 [species]
Onthophagus versutus Péringuey, 1901 [species]
Onthophagus verticalis Fahraeus, 1857 [species]
Onthophagus verticicornis Laicharting, 1781 [species]
Onthophagus vesanus Balthasar, 1967 [species]
Onthophagus vespertilio Howden & Cartwright & Hallfter, 1956 [species]
Onthophagus vestitus D'orbigny, 1902 [species]
Onthophagus vethi Krikken, 1977 [species]
Onthophagus vicinus Harold, 1886 [species]
Onthophagus victor Sharp, 1875 [species]
Onthophagus victoriensis Blackburn, 1903 [species]
Onthophagus viduus Harold, 1875 [species]
Onthophagus vigens Péringuey, 1901 [species]
Onthophagus vigilans Boucomont, 1921 [species]
Onthophagus vilis Harold, 1877 [species]
Onthophagus villanuevai Delgado & Deloya, 1990 [species]
Onthophagus villosus Macleay, 1888 [species]
Onthophagus vinctoides Frey, 1957 [species]
Onthophagus vinctus Erichson, 1843 [species]
Onthophagus violaceotinctus Gillet, 1925 [species]
Onthophagus violaceoviridis Paulian, 1937 [species]
Onthophagus violetae ZUNINO & HALFFTER, 1997 [species]
Onthophagus virens d'Orbigny, 1913 [species]
Onthophagus virescens Harold, 1867 [species]
Onthophagus viridiaereus D'orbigny, 1908 [species]
Onthophagus viridicatus D'orbigny, 1902 [species]
Onthophagus viridiceps d'Orbigny, 1910 [species]
Onthophagus viridicervicapra Ochi, Kon & Tsubaki, 2009 [species]
Onthophagus viridis Ménétriès, 1832 [species]
Onthophagus viriditinctus Reitter, 1893 [species]
Onthophagus viridivinosus Kohlmann & Solis, 2001 [species]
Onthophagus vitalisi Boucomont, 1921 [species]
Onthophagus vitulus Fabricius, 1776 [species]
Onthophagus vividus Arrow, 1907 [species]
Onthophagus viviensis D'orbigny, 1905 [species]
Onthophagus vladimiri Frey, 1957 [species]
Onthophagus vlasovi Medvedev, 1958 [species]
Onthophagus volucer Balthasar, 1959 [species]
Onthophagus vuattouxi Cambefort, 1984 [species]
Onthophagus vulpes Harold, 1877 [species]
Onthophagus vulpinaris Schönfeldt, 1906 [species]
Onthophagus vulpinus Arrow, 1931 [species]
Onthophagus vultuosus D'orbigny, 1902 [species]
Onthophagus vultur Arrow, 1931 [species]
Onthophagus vylderi D'orbigny, 1913 [species]
Onthophagus wagamen Matthews, 1972 [species]
Onthophagus wakelbura Matthews, 1972 [species]
Onthophagus wallacei Harold, 1871 [species]
Onthophagus walteri Macleay, 1887 [species]
Onthophagus waminda Matthews, 1972 [species]
Onthophagus wanappe Storey, 1977 [species]
Onthophagus wangi Masumoto, Chen & Ochi, 2004 [species]
Onthophagus wangnamkhieoensis Masumoto & Hanboonsong & Ochi, 2002 [species]
Onthophagus wangnamkieous Masumoto, Ochi & Hanboonsong, 2008 [species]
Onthophagus watanabei Ochi & Kon, 2002 [species]
Onthophagus waterhousei Gillet, 1927 [species]
Onthophagus waterloti D'orbigny, 1908 [species]
Onthophagus waterstradti Boucomont, 1914 [species]
Onthophagus watuwila Krikken & Huijbregts, 2008 [species]
Onthophagus wayaua Huijbregts & Krikken, 2012 [species]
Onthophagus wensis Josso & Prévost, 2006 [species]
Onthophagus weringerong STOREY & WEIR, 1990 [species]
Onthophagus werneri Moretto & Nicolas, 2002 [species]
Onthophagus wiebesi Krikken, 1977 [species]
Onthophagus wigmungan Matthews, 1972 [species]
Onthophagus wilgi Matthews, 1972 [species]
Onthophagus willameorum WALTER & CAMBEFORT, 1977 [species]
Onthophagus williamsi STOREY & WEIR, 1990 [species]
Onthophagus witteianus Krell, 2009 [species]
Onthophagus wombalano Matthews, 1972 [species]
Onthophagus worooa STOREY & WEIR, 1990 [species]
Onthophagus worsissa Roth, 1851 [species]
Onthophagus xanthochlorus WALTER & CAMBEFORT, 1977 [species]
Onthophagus xanthomerus Bates, 1887 [species]
Onthophagus xanthopterus D'orbigny, 1908 [species]
Onthophagus xanthopygus D'orbigny, 1908 [species]
Onthophagus xiphias Solis & Kohlmann, 2003 [species]
Onthophagus yackatoon STOREY & WEIR, 1990 [species]
Onthophagus yakuinsulanus Nakane, 1984 [species]
Onthophagus yamaokai Masumoto & Ochi & Hanboonsong, 2002 [species]
Onthophagus yangi Masumoto, Tsai & Ochi, 2006 [species]
Onthophagus yanoi Matsumura, 1938 [species]
Onthophagus yaran STOREY & WEIR, 1990 [species]
Onthophagus yarrumba Storey, 1977 [species]
Onthophagus yeyeko Matthews, 1972 [species]
Onthophagus yiryoront Matthews, 1972 [species]
Onthophagus yourula STOREY & WEIR, 1990 [species]
Onthophagus yubarinus Matsumura, 1937 [species]
Onthophagus yucatanus Delgado-Castillo, Peraza & Deloya, 2006 [species]
Onthophagus yukae Masumoto, Ochi & Hanboonsong, 2002 [species]
Onthophagus yumotoi Ochi & Kon, 2006 [species]
Onthophagus yungaburra Matthews, 1972 [species]
Onthophagus yunkara Matthews, 1972 [species]
Onthophagus yunnanus Boucomont, 1912 [species]
Onthophagus yvescambeforti Moretto, 2004 [species]
Onthophagus zagrosicus Kabakov, 2006 [species]
Onthophagus zairensis WALTER & CAMBEFORT, 1977 [species]
Onthophagus zapotecus ZUNINO & HALFFTER, 1988 [species]
Onthophagus zavattarii Müller, 1939 [species]
Onthophagus zavreli Balthasar, 1935 [species]
Onthophagus zebra Arrow, 1931 [species]
Onthophagus zebu Boucomont, 1921 [species]
Onthophagus zetteli Masumoto & Ochi & Hanboonsong, 2002 [species]
Onthophagus zicsii Balthasar, 1967 [species]
Onthophagus zimmermanni Balthasar, 1959 [species]
Onthophagus zumpti Frey, 1954 [species]
Onthophagus zuninoi Martin-Piera, 1985 [species]
Onthophagus zymoticus Moxey, 1963 [species]
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