Community and Forum → Insects biology and faunistics → Genus Carabus (Coleoptera, Carabidae) of Saratov region
Алексей Сажнев, 09.03.2010 14:05
MATERIALS ON THE DISTRIBUTION OF THE GENUS CARABUS LINNÉ, 1758 (COLEOPTERA, CARABIDAE) IN THE SARATOV REGION
A. S. Sazhnev
Saratov State Agrarian University named after N. I. Vavilov
on the rights of the manuscript
The classification was adopted according to B. Brezina (2004), taking into account the works of O. L. Kryzhanovsky, K. V. Makarov and co-authors (Kryzhanovsky, 1983; Kryzhanovskiy et. al., 1995; Makarov et al., 2007) and B. Lassele (2008). Synonymy is given by O. L. Kryzhanovskiy (Kryzhanovskiy et. al., 1995), V. G. Shilenkov (1996), B. Brezina (2004), and B. Lassele (2008).
Genus Carabus Linné, 1758
Typical view: granulatus Linné, 1758
Subgenus Carabus s. str.
[Carabus (s. str.) arcensis ssp. arcensis Herbst, 1784]
= arvensis Paykull, 1790; iss of arcensis Herbst, 1784
was previously considered as a possible species for the region (Sazhnev, 2007). The nominative subspecies is widely known on the territory of European Russia (Kryzhanovskij et al., 1995; Brezina, 2004), but its distribution relative to the Saratov region is not entirely clear; it is possible that the subspecies will be found in the north-west of the region
1. Carabus (s. str.) arcensis ssp. baschkiricus Breuning, 1936
Material: Saratov, 16. VI. 2008 (1) I. A. Zabaluev.
In Russia, the subspecies is distributed in the European part, mainly in the Volga region (Kryzhanovskij et al., 1995). From the region, the subspecies is known so far from one specimen from the vicinity of Saratov. It is found in various types of forests.
2. Carabus (s. str.) granulatus ssp. granulatus Linné, 1758
Material: Tatishchevsky district, Oktyabrsky Gorodok village, 18. V. 1896 (Sakharov, 1903: "Idolga River"); Tatishchevsky district, Vyazovka village, park, pod kamen, 2. VI. 1981 (1 female) O. A. Garanin (SOMK), ibid., 3.VI.1982 (1 male) O. A. Garanin (SOMK); Rtishchevsky district, Klyuchi village, "Makarovsky zakaznik", V. 1995 (3 specimens) I. V. Kryukov (Sazhnev, 2007); Voskresensky district, Usovka Village, 2004 (1 specimen) A. A. Kupriyanov (Sazhnev, 2007); Rtishchevsky district, Klyuchi village, "Makarovsky zapovednik", soil traps, 18. VI. 2007 (2 females, 1 male) I. V. Kryukov; Saratov region, collections of SSAU, 2007 (3 copies); Rtishchevsky district, Klyuchi village, "Makarovsky Zapovednik", 26-28. VI. 2008 (2 males) I. V. Kryukov, A. S. Sazhnev.
In the Saratov region, it is recorded only on the Right bank, however, the presence of the species in the Trans-Volga region is very likely in the north of the region.
It inhabits coastal habitats or moist floodplain forests and meadows with dense vegetation, occurs under river sediments in coastal talniki (Sakharov, 1903), in the south of the Volgograd region it is recorded in reed thickets along the banks of water bodies (Kalyuzhnaya et al., 2000).
3. Carabus (s. str.) stscheglowi Mannerheim, 1827
Material: Tatishchevsky district, Vyazovka village, on the litter as C. arvensis, 2. VI. 1977 (1 male) O. A. Garanin (SOMK), in the same place as C. arvensis, 17. VI. 1982 (1 male) O. A. Garanin (SOMK); Rtishchevsky district, S. Klyuchi, 29. V. 1993 (6 copies) I. V. Kryukov (Sazhnev, 2007); Saratov, 11. VI. 2005 (5 copies) (Sazhnev, Rodnev, 2005: "Lysogorsky massif"; Sazhnev, 2007); ibid.15.VI. 2005 (2 copies) (Sazhnev, 2007); Saratov, 5. VII. 2007 (1 female) A. S. Sazhnev; Saratov, 12. VII. 2007 (2 copies) N. V. Rodnev; Saratov, 11. V. 2007 (1 female, 1 male) A. S. Sazhnev; Saratov, 20. V.2007 (2 males, 2 females) A. S. Sazhnev; Saratov, 2. VII. 2007 (1 male) A. S. Sazhnev; Saratov, 5. VII. 2007 (1 female) A. S. Sazhnev; Saratov, 12. VII. 2007 (2 copies) N. V. Rodnev; Saratov, 3. VI. 2008 (1 male; 5 females) A. S. Sazhnev, N. V. Rodnev.
In the Saratov region, it is recorded only in the Right Bank: Saratovsky, Tatishchevsky, Rtishchevsky districts, but it is probably widespread everywhere: it has not been studied in the Trans-Volga region, but it is apparently absent on the border with Kazakhstan and in steppe and semi-desert zones. Every year it is common near Saratov (Lysogorsky massif, Kumysnaya Polyana).
It is noted in bayrachny, floodplain deciduous forests and plantings. In the conditions of the Volgograd region, it inhabits large gullies with massifs of bayrach forests of aspen, poplar, alder, and birch, usually occurring in damp, rocky places near streams and small rivers (E. V. Komarov).
Subgenus Limnocarabus Géhin, 1876
Type species: Carabus clathratus Linné, 1761
4. Carabus (Limnocarabus) clathratus ssp. clathratus Linné, 1761
Material: Krasnokutsky district, v sarae, 22. VI. 1988 (1 female) O. A. Garanin (SOMK); Balashovsky district, Nikolaevka village, deciduous forest, day, 20. V. 1989 (1 female) V. V. Anikin; Saratovsky district, Dubki village, state farm "Vesna", in teplitsa, 2. VII. 1997 (1 copy) I. V. Kryukov (probably brought together with peat from the Leningrad region) (Sazhnev, 2007); Saratov region, collection of SSAU, 2007 (1 copy).
In the literature (Kapranova et al., 2000), a species from the Dergachevsky and Fedorovsky districts is mentioned as a food element of the belladonna crane (Anthropoides virgo), however, a parallel reference is made to Carabus pannonicus L., a synonym of Carabus scheidleri ssp. scheidleri Panzer, 1799, which does not live in Russia, calls into question the accuracy of determining the species; in addition, the latter is listed under the Russian name as "Hungarian ground beetle" – and probably meant Carabus hungaricus scythus Motschulsky, 1847. The habitats of belladonna-steppe coenoses of the Saratov Trans-Volga region-also call into question the correctness of the determination of Carabus clathratus, known as a rather specialized hygrophilic species. Finally, the material was collected and reviewed fragmentally, since the study was performed on the feces of birds, not whole individuals, which also reduces the percentage of accuracy of the determination.
In the region, the species is widespread, apparently everywhere on the Right Bank and locally in the river valleys in the Trans-Volga region, so we leave the indications from the Dergachevsky and Fedorovsky districts as possible. Drive habitats, usually with dense vegetation. On clay or silted soil at the water's edge.
Subgenus Trachycarabus Géhin, 1885
Type species: Carabus scabriusculus Olivier, 1795
5. Carabus (Trachycarabus) estreicheri Fischer von Waldheim, 1822
= var. jaegeri Ménétriés, 1849
Material: Saratov region, without more precise indications, as " Carabus var.", collection of A. P. Kumakov, (COMK); Tatishchevsky district, S. Oktyabrsky gorodok, 12. IV. (Sakharov, 1903: "Nikolaevsky gorodok", as var. jaegeri Men.); Saratov, Oktyabrskoe gorge, 7. VI. 1988 (1 female) V. V. Anikin; Saratovsky district, Sinenkiye village, 21. VI. 1997 (1 specimen) I. V. Kryukov (Sazhnev, 2007); Saratov, 10-ya Dachnaya, VII.2003 (1 female) V. V. Anikin; Saratov, Lysogorsky massif, 6. V. 2005 (2 specimens) A. S. Sazhnev (Sazhnev, Rodnev, 2005); Saratov, 6. VI. 2005 (12 specimens) A. S. Sazhnev (Sazhnev, 2007); ibid.11.VI. 2005 (8 copies) A. S. Sazhnev, (Sazhnev, 2007); ibid. 15. VI. 2005 (9 copies) A. S. Sazhnev (Sazhnev, 2007); Saratov, Kumysnaya polyana, soil traps, 2007 (62 copies) A. S. Sazhnev; Saratov, Lysogorsky massif, 2nd Dachnaya, les, 11. V. 2007 (4 females) A. S. Sazhnev; ibid., 20. V. 2007 (3 females, 2 males) A. S. Sazhnev; ibid., 28.V.2007 (1 male) A. S. Sazhnev.
In the Saratov region, it is recorded only in the Saratov and Tatishchevsky districts. It is undoubtedly more widespread. Annually the dominant species in the vicinity of Saratov (Lysogorsky massif, Kumysnaya Polyana).
It is found in floodplain and bayrach forests, inhabits moist floodplain meadows and meadow grasses along the bottom of steppe gullies, and occurs under river sediments.
6. Carabus (Trachycarabus) sibiricus ssp. karelini Fischer von Waldheim, 1830
Material: Tatishchevsky district, village of Oktyabrsky gorodok (Sakharov, 1903-as Village of haeres Fisch., 1832); Saratov, in the park at the old cemetery, as Village of haeres, 6.VII.1986 (1 male) O. A. Garanin (SOMK); Saratov, sady, 19.V.1993 (1 copy) I. V. Kryukov (Sazhnev, 2007-as haeres Fisch., 1832); ibid., summer 1994 (1 copy) I. V. Kryukov (Sazhnev, 2007 – as haeres Fisch., 1832); Saratovsky district, Sinenkiye village, summer 1996 (1 copy) I. V. Kryukov (Sazhnev, 2007-as haeres Fisch., 1832). Kryukov (Sazhnev, 2007 - as haeres Fisch., 1832); ibid. 10. IX. 2006 (2 copies) I. V. Kryukov (Sazhnev, 2007 - as haeres Fisch., 1832); ibid., 1. III. 1997 (2 copies) I. V. Kryukov (Sazhnev, 2007 - as haeres Fisch., 1832); Saratov, 10. V. 1998 (1 copy) A. S. Sazhnev (Sazhnev, Rodnev, 2005 - as haeres Fisch., 1832; Sazhnev, 2007 - as haeres Fisch., 1832); Saratovsky district, O. p. Burkin, 3.VI. 2006 (1 copy) N. V. Rodnev (Sazhnev, 2007-as haeres Fisch., 1832); Saratov, pos. Uvek, 27. VII. 2008 (1 copy) SSAU fees.
In the Saratov region, it is recorded only in the Saratov and Tatishchevsky districts. It is undoubtedly more widespread. Early indications ofCarabus sibiricus haeres Fischer von Waldheim, 1823 for the Saratov region (Sakharov, 1903; Sazhnev and Rodnev, 2005; Sazhnev, 2007) were rechecked, and it was established that all specimens belong to C. sibiricus karelini Fischer von Waldheim, 1830. Unverified are the specimens based on those indicated by N. L. Sakharov (1903) – in the list of coleoptera for the Saratov province, these specimens are listed as " C. haeres Fisch.", however, the place of finds is" Nikolaevsky Gorodok", now St. Petersburg. Oktyabrsky gorodok, Tatishchevsky district, allows you to assign instances to ssp with a high degree of confidence. karelini Fischer von Waldheim, 1830.
It is noted in broad-leaved and mixed forests. N. L. Sakharov (1903) indicates beetles "in gardens under dry grass and various garbage".
Subgenus Archicarabus Seidlitz, 1887
Type species: Carabus nemoralis O. F. Muller, 1764
[Carabus (Archicarabus) nemoralis O. F. Muller, 1764]
Material: ?Saratov region, without more precise instructions, collection of A. P. Kumakov, (COMK) (possibly collections not from the Saratov region).
There is one copy of Carabus nemoralis from the collection of A. P. Kumakov in the SOMK funds, but the label was written later with the note"? Saratov region", and it is impossible to say for sure whether this material was collected from the territory of the Saratov region or not, this species was also previously recorded by us (Sazhnev and Rodnev, 2005), but it is doubtful that the material was also lost, so new finds of the species are needed. C. nemoralis is recorded from the territory of the Middle Volga region in adjacent areas. with the Saratov region in the same regions (Isaev et al., 2004), so we leave the view as possible for the region.
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