Georissidae Laporte, 1840

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Алексей Сажнев, 09.10.2011 13:23

GEORISSIDAE (COLEOPTERA: HYDROPHILOIDEA – - A NEW FAMILY OF COLEOPTERA FOR THE FAUNA OF THE SARATOV REGION

A. S. Sazhnev

Saratov State University named after N. G. Chernyshevsky

On the rights of the manuscript

Georissidae Laporte, 1840 – a small family of coleoptera, including only one genus Georissus Latreille, 1809, represented in all zoogeographical areas. According to S. V. Litovkin in the World Catalog (Hansen, 1999), the family includes 77 species. By 2010, at least 3 more species had been described (Fikáček & Trávníček, 2009; Makhan, 2009; Fikáček & Falamarzi, 2010). Thus, the family size is at least 80 species.

It should be noted that the fauna of the Palearctic region remained largely poorly understood. The Catalog of the Palearctic (Hansen, 2004) contains data on 19 species for the region; the list has been expanded to 21 species, taking into account the descriptions of Georissus (Neogeorissus) chameleo (Fikáček & Trávníček, 2009) and G. (N.) persicus (Fikáček & Falamarzi, 2010). Five species of Georissus are widely distributed in the European and Middle Eastern fauna: G. (s. str.) crenulatus (Rossi, 1794), G. (s. str.) substriatus Heer, 1841, G. (N.) caelatus Erichson, 1847, G. (N.) costatus Castelnau, 1840, and G. (N.) laesicollis Germar, 1831 (Fikáček & Falamarzi, 2010).

Geographical distribution of the genus Georissus.

По: Hebauer, F. 2004. Systematic and zoogeographical notes on the genus Georissus Latreille, 1809 (Coleoptera: Hydrophiloidea). Acta Coleopterologica 20 (1): 3–6.

There is no definitive list of Georissidae for Russia; according to various authors ' forecasts, up to 7 species of georissidae can be found in Russia (Zaitzev, 1910; Lafer, 1989; Kireichuk, 2001; Hansen, 2004).

No species of the genus Georissus were previously recorded in the Saratov region. From neighboring regions of the Volga region, there are data on the presence of G. (s. str.) crenulatus in Tatarstan (personal communication by D. A. Klemin), in the Ulyanovsk region (Isaev and Sysoenkov, 2000), and in the Samara region (Sachkov and Krasnobaev, 1998), probably based on unpublished materials by A.V. Burdaev, there are also modern ones findings of the species – personal report of S. V. Litovkin, G. (N.) costatus was also noted for the Samara region [Litovkin and Fikáček, in press].

This section will continue, comments, additions, links to sources or personal messages will be accepted (thank you all).

Larvae and adults are associated with different types of soil, live in the coastal zone of water bodies. Larvae are carnivorous, preying on small invertebrates such as fly larvae and nematodes (Hansen, 2000). Adult saprophages feed on decomposing organic residues (Hebauer, 1998). In some species, the upper side of the adult body is covered with a layer of silt or mud, which often complicates the search for beetles when collecting material. Adults of some species fly to the light.

The systematic position of this group has not been sufficiently elucidated and has changed several times, suggesting that it has many common features with Hydrophilidae (Crowson, 1981; Lawrence & Newton, 1995; Bernhard et al., 2009).

Currently, two collection points of Georissus are known for the Saratov region: in the Trans-Volga region-this is the Engelsky district of the region, and on the Right Bank-the territory of the Khvalynsky National Park. All the finds belong to Georissus (s. str.) crenulatus (Rossi, 1794). Instances are stored in collections of collectors.

Georissus (s. str.) crenulatus (Rossi, 1794)

Material:
Saratov region, Engels district, Engels village. Lesnoy, in silt on the shore of Lake Lesnoy, GPS: 51°29 '15.08" N 46°3 '40.64" E, 11. VII. 2010 (1 copy) I. A. Zabaluev leg., S. V. Litovkin, I. A. Zabaluev det., 2010; Saratov region, Khvalynsky district, Khvalynsky NP, on the shore of a forest lake, 7. VII. 2011 (1 copy) A. S. Sazhnev leg., A. S. Sazhnev det., 2011.

Photo from the site: http://www.colpolon.biol.uni.wroc.pl/geori...0crenulatus.htm

picture: Georyssus_crenulatus.jpg

Locale: Saratov region, Khvalynsky district, Khvalynsky NP, forest lake shore. Photo by A. S. Sazhnev

picture: _________.jpg

The author is grateful to I. A. Zabaluev (Saratov), S. V. Litovkin (Samara), and D. A. Klemin (Kazan) for their help and interest in the work. For assistance in conducting field research – V. V. Anikin (Saratov).

literature:

Isaev A. Yu., Sysoenkov D. A. 2000. K znaniyu vodolyuboobraznykh zhukov i vodoborok (Coleoptera: Hydrophiloidea: Hydrophilidae, Georyssidae; Staphylinoidea: Hydraenidae) Ul'yanovskoy oblasti [On the knowledge of water-loving beetles and water-bores (Coleoptera: Hydrophiloidea: Hydrophilidae, Georyssidae; Staphylinoidea: Hydraenidae) of the Ulyanovsk region]. Ulyanovsk. Issue 9. pp. 17-32.
Kireychuk A. G. 2001. Coleoptera (Coleoptera, or Beetles) / / In: Opredelitel ' presnovodnykh bespozvonochnykh Rossii i sopredel'nykh territorii. Volume 5. SPb, "Nauka", p. 91.
Lafer G. Sh. 1989. The family Georissidae. Determinant of insects of the Far East of the USSR (Volume III, Part 1). Leningrad, Nauka Publ., pp. 293-294.
Sachkov S. A., Krasnobaev Yu. P., 1998. Invertebrates of the Samara region: Handbook. Samara: Publishing House "Samara University". – 82 с.
Bernhard D., Ribera I., Komarek A., Beutel, R.G. 2009: Phylogenetic analysis of Hydrophiloidea (Coleoptera: Polyphaga) based on molecular data and morphological characters of adults and immature stages. Insect systematics & evolution, 40: 3–41.
Crowson R.A., 1981. The Biology of the Coleoptera. – Academic Press, London. P. 694–698.
Fikáček, M. & Falamarzi, Sh. 2010. Georissus persicus sp. nov. from Iran, with notes on the West-Palaearctic species of the G. laesicollis group (Coleoptera: Georissidae). Acta Entomologica Musei Nationalis Pragae 50: 107–116.
Fikáček, M. & Trávníček, D. 2009. Order Coleoptera, family Georissidae // Arthropod fauna of the UAE, 2: 145–148.
Hansen, M. 1999. World Catalogue of Isects. Vol. 2. Hydrophiloidea s. str. (Coleoptera). Apollo Books, Stenstrup, 416 pp.
Hansen, M. 2000. Observations on the immature stages of Georissidae (Coleoptera: Hydrophiloidea) with remarks on the evolution of the hydrophiloid egg cocoon. Invertebrate Taxonomy, 14: 907–916.
Hansen, M. 2004. Family Georissidae Laporte, 1840, p. 42. In: Löbl I. & Smetana A. (eds). Catalogue of Palaearctic Coleoptera, Vol. 2: Hydrophiloidea - Histeroidea - Staphylinoidea. Apollo Books, Steensrup, 942 pp.
Hebauer, F. 1998. Teil A: Imagines. Pp. 1–90 in: F. Hebauer, & B. Klausnitzer: Insecta: Coleoptera: Hydrophilidae (exkl. Helophorus). Süsswasserfauna von Mitteleuropa, 20, part 7, 8, 9, 10–1. Gustav Fischer, Stuttgart, Jena, Lübeck, Ulm, xii + 134 pp.
Lawrence J.F., Newton A.F., 1995. Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names) // In: Biology, Phylogeny, and Classification of Coleoptera. Eds. J. Pakaluk and S.A. Slipinski. Warszawa, 1995: 779–1006.
Litovkin, S.V., Fikáček, M. New records of Georissus costatus Laporte de Castelnau, 1840 (Coleoptera: Georissidae) from Russia // Russian Entomological Journal. (in print).
Makhan, D. 2009. Georissus amrishi sp. nov., a new water beetle from Suriname (Coleoptera: Georissidae). Calodema, 96: 1–5.
Zaitzev, P. 1910. Pars 17. Dryopidae, Cyathoceridae, Georyssidae, Heteroceridae. IN: S. Schenkling (ed.), Coleopterorum Catalogus. W. Junk, Berlin, 68 pp.

This post was edited by Alexey Sazhnev - 11.10.2011 12: 17

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10.10.2011 12:40, Алексей Сажнев

We are also currently translating the only version of the qualifier for European species. Currently, only the German version is available. We will always welcome the help of translators. If you can create a drawing key, you will need help with literature, drawings, and photo materials on five widely used types (see above). We hope that it will be useful not only for a narrow circle of professionals, but also for all lovers of insects and coleoptera.

Determinant of subgenera (according to Hebauer, 2004)

Hebauer, F. 2004. Systematic and zoogeographical notes on the genus Georissus Latreille, 1809 (Coleoptera: Hydrophiloidea). Acta Coleopterologica 20 (1): 3–6.

1. Pronotum is densely granular over the entire surface, the disc is uneven and weaker, the sides are without teeth, evenly curved. Elytra with alternating large and smaller keeled spaces, granular … Nipponogeorissus Satô, 1972
– - Pronotum slightly granular, or almost smooth on the disc ... 2.

2. Pronotum disc in a variety of pits, including a central (rarely narrowly elongated) and two oblique pits at the base of the pronotum, usually with more or less obvious grain, the sides often with lateral teeth in front of the posterior corners. The spaces of the elytra are usually granular, alternately raised ... Neogeorissus Satô, 1972
– - The pronotum disc (behind the anterior half) is uniformly convex, in extreme cases with a narrow central groove, smooth or with rare dots and grains, the sides without teeth. The spaces of the elytra are smooth, not raised … Georissus Latreille, 1809.

And the initial version of the determinant by type

based on:
Lafer G. S. 1989. The family Georissidae. Determinant of insects of the Far East of the USSR (Volume III, Part 1). Leningrad, Nauka Publ., pp. 293-294.
Steffan, A. W. 1979. 43. Familie: Georissidae. In: H. Freude, K. W. Harde and G. A. Lohse (eds.), Die Kaefer Mitteleuropas. Band 6. Diversicornia. Goecke & Evers, Krefeld. 294- 296.

illustrations in development

Genus: Georissus Latreille 1809
(Georyssus Agassiz, 1846, misspelled)

The only genus of the family. In Central Europe, it is represented by 5 species. The species are entirely black, with family characteristics.

1. The surface of the pronotum is sculptured only in the anterior half, the posterior part is largely smooth. Elytra with simple rows of dots ... 2 (Subgenus Georissus Latreille 1809).
- The entire surface of the pronotum is roughly sculptured. Elytra ribbed or grooved ... 3. (Subgenus Neogeorissus Satô, 1972)

Photo materials from the site: http://www.iran-eng.com/showthread.php/193...leoptera/page28 - the author of the photo is K. V. Makarov.

picture: 1.jpg

explanations in the text

Subgenus Georissus Latreille 1809

2. The anterior half of the pronotum is roughly sculptured, with a deep groove at the anterior edge and along the sides, as well as a deep longitudinal groove in the middle of the anterior half of the pronotum. The posterior half of the disc is convex and smooth. Elytra with longitudinal rows of round large and deep dots, row spacing slightly convex, smooth and wide, approximately equal to the diameter of the dots in the rows. The upper body is shiny. Black. 1.5-2.1 mm ... crenulatus (Rossi, 1794).

- Pronotum in the anterior part is weakly or practically not sculptured, only grainy shagreen. Longitudinal rows of elytra with much less deep, shallow, sometimes disappearing points, the rows themselves are located at a greater distance from each other. The upper part of the body is matte. Black. 1.6-1.8 mm ... substriatus Heer, 1841.

Из: E. Reitter "Die Kaefer des Deutschen Reiches", Band: III. Tab. 104. 1911.

Image: Georissus. png

Georissus substriatus Heer, 1841.
Иллюстрация из: Merkl O. 2006: New beetle species in the Hungarian fauna (Coleoptera) – Folia entomologica hungarica 67: 19-36.

image: substriatus. png